New taxa of the Lygistorrhinidae (Diptera: Sciaroidea) and their implications for a phylogenetic analysis of the family Author Hippa, Heikki Author Mattsson, Ingegerd Author Vilkamaa, Pekka text Zootaxa 2005 960 1 34 journal article 10.5281/zenodo.171227 99e6920b-84b2-497c-9b7b-5421aa657a5c 1175­5326 171227 Key to genera of Lygistorrhinidae We include Lygistorrhina asiatica in the key separately because a new genus is needed for it. According to the phylogenetic analysis ( Fig. 1 ) it is the sister group of all other living Lygistorrhinidae . Lygistorrhina ( Lygistorrhina ) and Lygistorrhina ( Probolaeus ) are separated even if their monophyly is not yet shown. The sexual dimorphism in the species of the Lygistorrhinidae is largely unknown and it is possible that some of the characters used below apply only to males. 1. Number of ocelli 3......................................................................................................... 2 ­ Number of ocelli 2......................................................................................................... 6 2. Rs present, sclerotized............................................................. Palaeognoriste Meunier ­ Rs absent or discernible only as a non­sclerotized fold ............................................... 3 3. Proboscis short, at most two and a half times the height of head ................................ 4 ­ Proboscis long, four or more times as long as the height of head ................................ 5 4. Subcosta ending in costa, R1 meeting costa at the middle of wing ............................... ...........................................................................” Lygistorrhina ” asiatica Senior­White ­ Sc ending free, R1 meeting costa well within the basal half of wing ............................ ..................................................................................................... Labellorrhina gen. n. 5. Middle tibia with two apical spurs ............. Lygistorrhina Skuse , subg. Lygistorrhina ­ Middle tibia with one apical spur.... Lygistorrhina Skuse , subg. Probolaeus Williston 6. Hind basitarsomere swollen, as broad as apical part of hind tibia ............................... 7 ­ Hind basitarsomere thin, narrower than apical part of hind tibia ( Figs 5 B, 6B, 7B) .... 9 7. C with a swelling at the apex of vein R1, proboscis very short, about one­tenth the height of head ...................................................................................... Seguyola Matile ­ C without a swelling at the apex of R1, proboscis long, as long as or longer than the height of head ............................................................................................................... 8 8. Number of antennal flagellomeres 12, R1 meeting the costal margin well in basal half of wing .................................................................... Loyugesa Grimaldi & Blagoderov ­ Number of antennal flagellomeres14, R1 meeting the costal margin at the middle of wing Matileola Papp