The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae) Author Wagner, Rüdiger Author Withers, Phil text Zootaxa 2020 2020-01-22 4728 2 183 210 journal article 24286 10.11646/zootaxa.4728.2.2 a1c40b73-0c11-4863-a188-412c33523e02 1175-5326 3621117 AE4F8089-7D2E-4ABA-AA09-CD504705788A Key to males of the west-palearctic species of genus Tonnoiriella Vaillant 1982 1 - Hypandrium thin without broadening, gonostyli 1.5 x longer than gonocoxites with angled tip, aedeagus with parallel sides, longer than gonocoxites, inner seta on gonocoxites shorter than the diameter of the gonocoxite...... T. graeca 1993 Wagner - Hypandrium usually with median broadening, gonostyli about as long as gonocoxites with sharp tip, gonocoxites with a strong seta long distinctly longer than the gonocoxite diameter....................................................... 2 2 - Dorsal lobus of the basiphallus linked with a clearly detectable distiphallus sclerite................................. 3 - Dorsal lobus of the basiphallus without detectable distiphallus sclerite.......................................... 12 3 - Dorsal distiphallus extensive, lateral protrusions with sharp tips................................ T. pulchra -group ... 4 - Dorsal distiphallus small anchor-shaped, not protruding the edge of the aedeagus................... T. sieberti -group … 7 4 - Hypandrium medially with a thin tongue-shaped broadening................................. T. anderssoni (Nielsen) - Hypandrium medially with large broadening................................................................ 5 5 - Median broadening of hypandrium concave..................................................... T. paveli Ježek - Median broadening of hypandrium convex................................................................. 6 6 - Median broadening of hypandrium rhomboid................................................. T. pulchra (Eaton) - Median broadening of hypandrium triangular................................................ T. aurasica sp. nov. 7 - Hypandrium thin, of equal width over the entire length.......................................... T. syriensis Ježek - Hypandrium distinctly broadened in the midst.............................................................. 8 8 - Hypandrium broadened in the midst and less pronounced laterally............................................... 9 - Hypandrium broad over the entire ventral length, with distinct central broadening................................. 11 9 - Hypandrium broad only in the middle........................................................ T. sieberti Wagner - Hypandrium with in addition 2 smaller lateral prominences................................................... 10 10 - Ventral distiphallus sclerite as long as gonocoxite, apically with 2 small tips.......................... T. ikariae sp. nov. - Ventral distiphallus sclerite shorter than gonocoxite, apically blunt................................. T. turcica Wagner 11 - Hypandrium ventrally only slightly broadened................................................. T. egeica sp. nov. - Hypandrium ventrally in the midst shield-like broadened......................................... T. italiae sp. nov. 12 - Dorsal lobus of basiphallus apically dualistic branched....................................................... 13 - Dorsal lobus of basiphallus not branched.................................................................. 14 13 - Median broadening of hypandrium 2x longer than broad...................................... T. atlantica (Satchell) - Median broadening of hypandrium inconspicuous................................. T. fontinalis Wagner & Salamanna 14 - Gonostylus apically apruptly thinner, digitate or harpoon shaped with serrate outer edge............. T. mollis -group … 15 - Gonostylus evenly bent, tapered, epandrial processes appendages elongate with more than 5 retinacula................................................................................................. T. nigricauda -group … 17 15 - Hypandrium medially hardly broadened, epandrial processes appendages with 1 large and 2 short tenacula.................................................................................................... T. mollis (Satchell) - Hypandrium medially linguiform broadened............................................................... 16 16 - Gonostylus as long as gonocoxite, basally globular, apically harpoon-shaped............... T. anchoriformis (Salamanna) - Gonostylus longer than the gonocoxite, apically digitate with a line of 4 setae bases........ T. filistylis Vaillant & Moubayed 17 - Hypandrium with 2 median or lateral broadenings.......................................................... 18 - Hypandrium with 1 median broadening................................................................... 19 18 - Broadenings lateral, short, acute, ventral distiphallus sclerite almost semicircular apically acute........... T. arcuata Ježek - Broadenings close to the middle, rounded, ventral distiphallus sclerite apically round............... T. goncalvesi sp. nov. 19 - Median broadening small, ventral distiphallus sclerite with blunt 3 lobes....................... T. nigricauda (Tonnoir) - Median broadening distinct, ventral distiphallus sclerite with 2 sharp lobes......................... T. andradei sp. nov.