The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama Author Blahnik, Roger J. Author Armitage, Brian J. text Insecta Mundi 2019 2019-12-23 740 740 1 17 journal article 10.5281/zenodo.3676607 5fb7c238-0813-4cf2-9c99-b827f92c398d 1942-1354 3676607 B942B8CA-4EEE-4C7A-A4B7-987C629E9DDA Mortoniella ( Mortoniella ) calovebora Blahnik and Armitage , new species Fig. 2 Diagnosis. This is a rather typical, but distinctive, species of the “florica subgroup.” Mortoniella calovebora is easily distinguished from other described species of the subgroup by the length and asymmetry of the apicoventral lobes of the inferior appendages ( Fig. 2C ). The asymmetry of the ventral lobes of the inferior appendages seems to be a consistent feature of this species and is a unique character within the genus. Description. Length of forewing: male 3.0– 3.3 mm ; female 3.2–3.3 mm . Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0:4:4. Overall color (in alcohol) light brown, anastamosis unpigmented, but otherwise not evidently marked. Table 1. List of species in the Glossosomatidae from Panama. The literature citation in which each species was first mentioned from Panama is given, if different from that of an original description (*) or this paper (+).

Species	Initial Literature Citation	Distribution
Suborder Integripalpia Martynov
Superfamily Glossosomatoidea Wallengren
Family Glossosomatidae Wallengren
Culoptila costaricensis Flint, 1974	+	Costa Rica*, Panama
Culoptila unispina Blahnik and Holzenthal, 2006	*	Costa Rica*, Panama
Mortoniella akantha Blahnik and Holzenthal, 2008	Armitage et al. 2018	Costa Rica*, Panama
Mortoniella anakantha Blahnik and Holzenthal, 2008	*	Costa Rica*, Panama
Mortoniella aviceps Blahnik and Holzenthal, 2008	*	Costa Rica*, Panama
Mortoniella calovebora Blahnik and Armitage, 2019	+	Panama*
Mortoniella carinula Blahnik and Holzenthal, 2008	Armitage et al. 2016	Costa Rica*, Panama
Mortoniella falcicula Blahnik and Holzenthal, 2008	Armitage et al. 2018	Mexico*, Panama
Mortoniella munozi Blahnik and Holzenthal, 2008	*	Costa Rica*, Panama
Mortoniella opinionis Blahnik and Holzenthal, 2008	+	Costa Rica*, Panama
Mortoniella panamensis Blahnik and Holzenthal, 2008	*	Panama*
Mortoniella pectinella Blahnik and Holzenthal, 2008	*	Panama*
Mortoniella redunca Blahnik and Holzenthal, 2008	Armitage et al. 2015	Costa Rica*, Panama
Mortoniella rovira (Flint, 1974)	*	Costa Rica, Panama*
Mortoniella tapanti Blahnik and Holzenthal, 2008	*	Costa Rica*, Panama
Mortoniella taurina Blahnik and Holzenthal, 2008	*	Costa Rica*, Panama
Mortoniella umbonata Blahnik and Holzenthal, 2008	*	Panama*
Mortoniella yayas Blahnik and Armitage, 2019	+	Panama*
Protoptila altura Holzenthal and Blahnik, 2006	Armitage et al. 2015	Costa Rica*, Panama
Protoptila boruca Flint, 1974	Armitage et al. 2018	Costa Rica*, Panama
Protoptila cana Flint, 1974	*	Costa Rica*, Panama
Protoptila chitaria Holzenthal and Blahnik, 2006	Armitage et al. 2015	Costa Rica*, Panama
Protoptila inflata Blahnik and Armitage, 2019	+	Panama*
Protoptila laterospina Flint, 1967	McElravy et al. 1981	Costa Rica*, Panama
Protoptila orotina Flint, 1974	*	Costa Rica*, Panama
Protoptila perdida Bueno-Soria, Santiago-Fragosa, and Barba-Alvarez, 2004	*	Panama*
Protoptila rambala Blahnik and Armitage, 2019	+	Panama*
Protoptila spirifera Flint, 1974	+	Costa Rica*, Panama
Protoptila tojana Mosely, 1954	Holzenthal and Blahnik 2006	Costa Rica, Honduras, Mexico*, Nicaragua, Panama, Peru
Protoptila totumas Blahnik and Armitage, 2019	+	Panama*
Protoptila trichoglossa Holzenthal and Blahnik, 2006	Armitage et al. 2015	Costa Rica*, Panama

Male genitalia. Ventral process of segment VII laterally compressed, short, ventrally projecting, rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded dorsally, narrowing ventrally; segment deeply excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively elongate, lateral margins subparallel, apically with short, subtriangular mesal projection; apicolateral lobes elongate, compressed, subacute, slightly mesally curved; ventrolateral lobes nearly obsolete; ventromesal lobes not evident. Inferior appendages without dorsolateral lobes, but with apicoventral lobes elongate, upturned, and asymmetrically developed, that of the right side narrower apically and more distinctly out-turned, both lobes acute apically. Mesal pockets of inferior appendage with apical processes short and weakly curved. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, slightly widened preapically, widened apex with narrow, adpressed, scale-like spines. Dorsal phallic spine, as viewed laterally, undulate in contour, spine distinctly widened on ventral margin at about middle, ventral bulge articulating with dorsal margin of phallicata, apex acute, weakly upturned; in dorsal view ( Fig. 2D ), lightly widened in middle and preapically, apex acute. Phallicata moderately elongate, dorsal margin with broadly rounded, paired, dorsolateral projections, lateral margin with slightly produced, rounded and compressed projection. Endophallic membrane simple, with single prominent curved ventromesal spine; phallotremal spines absent. Female and immatures. Unknown. Holotype male (alcohol). PANAMA : Veraguas : Cuenca 097, Río Calovebora , 8.55038°N and 81.16486°W , PSPSCD-PNSF-C097-2017-006, 461 m , Malaise trap , 23–27.iv.2017 , A. Cornejo , T . Ríos, E . Álvarez, and C. Nieto , ( COZEM ). Paratypes . PANAMA : Veraguas : Cuenca 097, Río Calovebora , 8.55038°N and 81.16486°W , PSPSCD- PNSF-C097-2017 - 006 , 461 m , Malaise trap , 23–27.iv.2017 , A. Cornejo , T . Ríos, E . Álvarez, and C. Nieto , 7 males ( COZEM ) ; ibid., Cuenca 132, Río Mulaba–Isleta , 8.54513°N and 81.1197°W , PSPSCD-PNSF- C132-2017-015, 412 m , Malaise trap , 2 males and 4 females ( COZEM ) ; ibid., Río Mulaba–afluente 1 er Brazo , 8.51706°N and 81.1214°W , Control : PSPSCD-PNSF-C132-2017-008, 770 m , UV light trap, 19.iv.2017 , T . Ríos, E . Álvarez, and C. Nieto , 1 male ( UMSP ) . Etymology. This new species is named for the Río Calovebora from which it was first collected. The name is formulated as a noun in apposition.