Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography Author Merle, Didier Author Pacaud, Jean-Michel Author Métais, Grégoire Author Bartolini, Annachiara Author Lashari, Rafiq A. Author Brohi, Imdad A. Author Solangi, Sarfraz H. Author Marivaux, Laurent Author Welcomme, Jean-Loup text Zootaxa 2014 3826 1 101 138 journal article 45364 10.11646/zootaxa.3826.1.3 6f4b1535-6c6b-4446-aa26-b1ba83ad9126 1175-5326 228537 53549D58-8F38-47B5-879F-0245E900C131 Athleta (Volutospina) noetlingi ( Cossmann & Pissarro, 1909 ) Volutospina noetlingi Cossmann & Pissarro, 1909 : 24 , pl. 2, figs 23–24. ? Voluta cithara (non Voluta cithara Lamarck, 1803 ). d’Archiac & Haime, 1853: 325, pl. 32, figs 4–5. Athleta (Neoathleta) noetlingi . Vredenburg 1923 : 256; Cotter in Vredenburg 1928 : 36 [new combination]. Lyria cossmanni (non Lyria cossmanni Vredenburg, 1923 ). Iqbal 1972 : 65, pl. 19, figs 3, 9. Comments. According to Cossmann & Pissarro (1909) , the specimen illustrated on their pl. 2, figs 23–24 displays a biconic shape with spiny costae delineating a developed shoulder and looks more like a species of Volutospina than a species of Volutopupa Dall, 1890 ( type species: Voluta cithara Lamarck, 1803 by original designation). We designate here this specimen as the lectotype of V. noetlingi . Vredenburg (1923) regarded Volutospina intercrenata Cossmann & Pissarro, 1909 as a junior synonym of V. noetlingi , considering that the specimens used to describe V. intercrenata represent juvenile specimens of V. noetlingi . However, the figured material of V. intercrenata does not display spiny costae delineating a developed shoulder, as observed in juvenile specimens of Volutospina species (e.g. the type species). Thus, these specimens do not belong to Volutospina and are not juveniles of V. noetlingi . In the Lakhra Dome, we have not found specimens of V. noetlingi , but a juvenile specimen of A. (V.) noetlingi from Jhirak has been illustrated by Iqbal (1972) under the erroneous name Lyria cossmanni . As in other juvenile specimens of Volutospina species, it has a biconic shape with spiny costae delineating a developed shoulder. Stratigraphic range. Lakhra Formation: Jhirak ( Cossmann & Pissarro 1909 ; Vredenburg 1923 ).