Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae) Author Lin, Shou-Wang Author Lopardo, Lara Author Uhl, Gabriele text Zoological Journal of the Linnean Society 2022 2021-11-20 195 417 584 journal article 115951 10.1093/zoolinnean/zlab033 75b10efe-a17d-4d10-b46e-094c6a7ebf01 0024-4082 6967774 BE2B3859-8F6A-4543-8A69-91840F82377B HOLMELGONIA BASALIS JOCQUÉ & SCHARFF, 1986 ( FIGS 19C , 22C , 24C , 36 ; SUPPORTING INFORMATION, FIG. S2A ) Elgonella basalis Jocqué & Scharff, 1986: 12 , figs 18–25 (Dmf). Elgonia basalis Platnick, 1989: 232 . Elgonia basalis Scharff, 1990a: 25 , figs 44–46 (mf). Holmelgonia basalis , Jocqué & Scharff, 2007: 161 . Examined material: Tanzania , Uzungwa Mts. , Mwanihana Forest above Sanje River , 1700 m , 1♂ 1♀ 15.viii.1982 , leg. M, Stoltze & N, Scharff ( ZMUC ) ; same locality, 1800 m , pitfall trap , 1♂ 18.viii.1982 , leg. M, Stoltze & N, Scharff ( ZMUC ) ; same locality, 1800 m , litter, 1♀ 15.viii.1982 , leg. M, Stoltze & N, Scharff ( ZMUC ) . Description: Male ( ZMUC , 15.viii.1982 ): Total length: 1.72. Prosoma: 0.79 long, 0.61 wide, without modification ( Fig. 19C ). Eyes: AME-AME: 0.02, AME width: 0.04, AME-ALE: 0.02, ALE width: 0.08, ALE-PLE: 0, PLE width: 0.09, PLE-PME: 0.08, PME width: 0.02, PME- PME: 0.03. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.44 long, 0.48 wide. Chelicerae: mastidia absent; stridulatory striae ridged, rows widely and evenly spaced ( Fig. 22C ). Legs: tibia chaetotaxy 2-2-1-1; dorsal proximal macroseta on tibia II, III and IV 2.24, 4.1 and 3.85 times diameter of tibia, respectively; Tm I: 0.60. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPA large, base distant from C base, distal part as wide as base, with two protuberances at tip, each with a seta below ( Fig. 36A, C ); PC base not visible from dorsal view, distal setae close to distal clasp, distal-setae-bearing area wide, distal clasp without striae, clasp extended apically ( Fig. 36A ); T without papillae, PT apical part with long papillae; TS shot, slender ( Fig. 36D ); MSA present; DSA wide, truncated at tip ( Fig. 36A ); EM broad, flat, without papillae, not exceeding ARP; ARP pointed; LER absent; central embolar apophysis present; VRP long, with papillae close to tip; TP long, round at tip; E short and slender, slightly retrolaterally curved ( Fig. 36B ). Opisthosoma: dorsal pattern see Fig. 24C ; PMS with mAP, AC absent; PLS with triad, one AC ( Fig. 36G ). Figure 36. Holmelgonia basalis Jocqué & Scharff, 1986 . A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets. H, female right spinnerets. I, female right posterior lateral spinneret. Scale bars 0.1 mm. Female: Total length: 2.04. Prosoma: 0.88 long, 0.62 wide. Eyes: AME-AME: 0.02, AME width: 0.04, AME-ALE: 0.02, ALE width: 0.08, ALE-PLE: 0, PLE width: 0.07, PLE-PME: 0.04, PME width: 0.07, PME- PME: 0.03. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.5 long, 0.51 wide. Legs: tibia chaetotaxy 2-2-1-1; dorsal proximal macroseta on tibia I 2.85 times diameter of tibia, respectively; Tm I: 0.60. Epigyne: Clade 13 characteristic morphology, CO close to dorsal plate posterior margin ( Fig. 36E, F ). Opisthosoma dorsal pattern same as male; PMS with mAP, CY; PLS with triad, one AC, two CY ( Fig. 36H, I ). Distribution: Tanzania .