Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species Author Grego, Jozef Horna Micina 219, 97401, Banska Bystrica, Slovakia Author Mumladze, Levan Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia https://orcid.org/0000-0002-2172-6973 Author Falniowski, Andrzej Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland Author Osikowski, Artur Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland https://orcid.org/0000-0001-6646-2687 a.osikowski@urk.edu.pl Author Rysiewska, Aleksandra Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland Author Palatov, Dimitry M. Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation Author Hofman, Sebastian Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland text ZooKeys 2020 955 1 77 http://dx.doi.org/10.3897/zookeys.955.51983 journal article http://dx.doi.org/10.3897/zookeys.955.51983 1313-2970-955-1 C8EF4A4B6F054621B9B5AE47FEE7C217 F7018CF19C6158309D57C83A2F5E9478 Kartvelobia sinuata Grego & Mumladze sp. nov. Plates 1 (1-3); 2(1-15); 3(1-4); 4(5-7); 5(3); Fig. 9A-E Type locality. Georgia • Imereti, Satsiskvilo, Turchusmtha (თურჩუსმთა, სოფელი საწისქვილო ); 42°29'49"N , 42°32'49"E ; 980 m a.s.l.; Small spring at left side of path to the plateau. Material. Holotype : Georgia • 1 adult, dry; type locality; 02 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg.; ISU FM-T020-H. Paratypes : Georgia • same as for holotype; ISU FM-T020-P1/500 dry specimens, SBMNH 633041/7 dry, NHMW 113374/10 dry, HNHM 104683/10 dry, NHMUK 20191339/10 dry, NHMB 563971/10 dry, SMF 358930/10 dry, ZMH 140635/10 dry, MNHN-IM-2016-7894, ZIN 1/511-2020/10 dry, coll. JG F0989/500 dry, coll. Gloeer /2 dry; Other material : Georgia • Imereti, Nakhriduri, Turchu Gamosadivari Cave; 42°29'13"N , 42°31'20"E ; 951 m a.s.l.; 03 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg., JG/3 dry. • Imereti, Nakhriduri 3 spring at Turchu Gamosadivari Basin left side; 42°28'41"N , 42°30'45"E ; 860 m a.s.l.; 03 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg. JG/5 dry. • Imereti, Nakhriduri 2, Turchu Gamosadivari Basin left side spring above small ford; 42°28'39"N , 42°30 43"E ; 860 m a.s.l.; 03 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg., JG/6 dry. • Imereti, Kinchkhaperdi (კინჩხაფერდი), spring right along the road to Askhi Plateau; 42°30'7"N , 42°33'34"E ; 880 m a.s.l.; 02 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg. JG/3 dry. • Imereti, Upskhero Spring Lake (უფსკერო ტბა ) at Turchu Gamosadivari Basin; 42°27'47"N , 42°30'3"E ; 890 m a.s.l.; 03 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg. JG/6 dry. • Imereti, Turchusmtha, spring of Okatse above Kinchkha waterfall; 42°29'49"N , 42°32'49"E ; 1050 m a.s.l.; 02 May 2018; J. Grego, L. Mumladze and M. Olsavsky leg. JG/5 dry. Plate 1. 1-3 Kartvelobia sinuata sp. nov. Imereti, Satsiskvilo, Turchusmtha, specimens used for molecular and anatomical study. Measurement of aperture declination angle (CA): aperture-elongation axis vs. the columella is depicted in green. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph A. Falniowski. Plate 2. 1-5 Kartvelobia sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha 1 holotype 2-5 paratypes 6 K. sinuata sp. nov., Imereti, Turchusmtha, spring of Okatse above Kinchkha waterfall 7, 8 K. cf. sinuata , Samegrelo, Pirveli Balda, Motena Cave 9-11 K. cf. sinuata , Samegrelo, Pirveli Balda, spring in village 12 K. sinuata sp. nov., Imereti, Nakhriduri, spring cave Turchu Gamosadivari 13 K. sinuata sp. nov., Imereti, Nakhriduri spring at Turchu Gamosadivari Basin 14, 15 K. sinuata sp. nov., Imereti, Nakhriduri side spring at Turchu Gamosadivari Basin. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego. Plate 3. Kartvelobia sinuata sp. nov. 1 Imereti, Turchu Gamosadivari Basin, Nakiduri spring, paratype SBMNH 633100 2 K. sinuata sp. nov., Imereti, Turchu Gamosadivari Basin, Nakhriduri 5, paratype SBMNH 633103 3 K. sinuata sp. nov., Samegrelo, Pirveli Balda, Motena Cave, paratype SBMNH 633043 4 K. sinuata sp. nov., Samegrelo, Pirveli Balda, spring in village, paratype SBMNH 633070. SEM SBMNH Vanessa Delnavaz. Scale bars: 1 mm.