<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1017.34500" ID-GBIF-Dataset="878575d9-dbd8-4d55-90ee-f8572e71cb81" ID-PMC="PMC7895820" ID-Pensoft-Pub="1313-2970-1017-37" ID-Pensoft-UUID="68184E5B74245C1F83F25E846E5608F6" ID-PubMed="33633482" ID-ZooBank="390FEA39DEBA4406B99FBC6625821960" ModsDocID="1313-2970-1017-37" checkinTime="1613177915340" checkinUser="pensoft" docAuthor="Acal, Dale Ann P., Wiesner, Juergen, Nuneza, Olga M. &amp; Jaskula, Radomir" docDate="2021" docId="9E71DEBD0DE257F1ADD8798C7047898C" docLanguage="en" docName="ZooKeys 1017: 37-75" docOrigin="ZooKeys 1017" docPubDate="2021-02-12" docSource="http://dx.doi.org/10.3897/zookeys.1017.34500" docTitle="Therates coracinus subsp. coracinus coracinus Erichson 1834" docType="treatment" docVersion="4" id="68184E5B74245C1F83F25E846E5608F6" lastPageNumber="37" masterDocId="68184E5B74245C1F83F25E846E5608F6" masterDocTitle="Tiger beetles (Coleoptera, Cicindelidae) of Northern Mindanao region (Philippines): checklist, distributional maps, and habitats" masterLastPageNumber="75" masterPageNumber="37" pageNumber="37" updateTime="1668149799970" updateUser="ExternalLinkService">
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<mods:title>Tiger beetles (Coleoptera, Cicindelidae) of Northern Mindanao region (Philippines): checklist, distributional maps, and habitats</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Acal, Dale Ann P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8102-5116</mods:nameIdentifier>
<mods:affiliation>Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Andres Bonifacio Ave., Tibanga, Iligan City 9200, Philippines</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiesner, Juergen</mods:namePart>
<mods:affiliation>Dresdener Ring 11, D- 38444, Wolfsburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nuneza, Olga M.</mods:namePart>
<mods:affiliation>Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Andres Bonifacio Ave., Tibanga, Iligan City 9200, Philippines</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jaskula, Radomir</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8949-848X</mods:nameIdentifier>
<mods:affiliation>Department of Invertebrate Zoology and Hydrobiology, Faculty of Biology and Environmental Protection, University of Lodz, Banacha 12 / 16, 90 - 237, Lodz, Poland</mods:affiliation>
<mods:nameIdentifier type="email">radomir.jaskula@biol.uni.lodz.pl</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-12</mods:number>
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<mods:detail type="volume">
<mods:number>1017</mods:number>
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<mods:start>37</mods:start>
<mods:end>75</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1017.34500</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1017.34500</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1017-37</mods:identifier>
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<treatment ID-GBIF-Taxon="177383223" LSID="urn:lsid:plazi:treatment:9E71DEBD0DE257F1ADD8798C7047898C" httpUri="http://treatment.plazi.org/id/9E71DEBD0DE257F1ADD8798C7047898C" lastPageNumber="37" pageId="0" pageNumber="37">
<subSubSection pageId="0" pageNumber="37" type="nomenclature">
<paragraph pageId="0" pageNumber="37">
<taxonomicName LSID="9E71DEBD-0DE2-57F1-ADD8-798C7047898C" authority="coracinus Erichson, 1834" authorityName="coracinus Erichson" authorityYear="1834" class="Insecta" family="Cicindelidae" genus="Therates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Therates coracinus subsp. coracinus" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="coracinus" subSpecies="coracinus">Therates coracinus coracinus Erichson, 1834</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitats of tiger beetles from Northern Mindanao region: Calomera angulata angulata (C-E) (sites 10 - 12), C. lacrymosa (C, E) (sites 10, 12), C. mindanaoensis (C-F, H) (sites 10 - 13, 15), Cylindera discreta elaphroides (A, C-E) (sites 9 - 12), C. minuta (A, C-E) (sites 9 - 12), C. viduata (G) (site 14), Lophyra striolata tenuiscripta (G) (site 14), Therates coracinus coracinus (B) (site 10) (descriptions of sites in Table 1; photographs A, C-H DAPA, B RJ)." figureDoi="10.3897/zookeys.1017.34500.figure3" httpUri="https://binary.pensoft.net/fig/509189" pageId="0" pageNumber="37">Figures 3B</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitats of tiger beetles from Northern Mindanao region: Calomera mindanaoensis (C) (site 16) (E, F) (sites 18 - 19), Prothyma heteromallicollis heteromallicollis (A) (site 16), Neocollyris similior (D) (site 17), Therates coracinus coracinus (D) (site 17), T. fasciatus fasciatus (D) (site 17), and Tricondyla aptera punctipennis (B) (site 16) (descriptions of sites in Table 1; photographs A-C RJ, D O. Bagona, E A. B. Lapore, F C. Torres)." figureDoi="10.3897/zookeys.1017.34500.figure4" httpUri="https://binary.pensoft.net/fig/509190" pageId="0" pageNumber="37">, 4D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of A Therates coracinus coracinus B Therates fasciatus fasciatus (circle) and T. fasciatus pseudolatreillei (square) C Therates fulvipennis bidentatus (cirle) and T. fulvipennis everetti (square) D Prothyma (Symplecthyma) heteromallicollis heteromallicollis E Heptodonta nigrosericea, and F Calomera angulata angulata in Northern Mindanao region." figureDoi="10.3897/zookeys.1017.34500.figure8" httpUri="https://binary.pensoft.net/fig/509194" pageId="0" pageNumber="37">, 8A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="distribution">
<paragraph pageId="0" pageNumber="37">General distribution.</paragraph>
<paragraph pageId="0" pageNumber="37">
Subspecies known from Indonesia, Moluccas, and Philippines, where it was recorded in Balabac, Leyte, Luzon, Mindanao, Mindoro, Negros, Palawan, Panay, Romblon, and Samar; in Mindanao recorded from Davao, Northern Mindanao, and Soccsksargen regions (Wiesner 1988; 
<bibRefCitation author="Cabras, AA" journalOrPublisher="Lambillionea" pageId="0" pageNumber="37" pagination="188 - 201" refId="B6" refString="Cabras, AA, Cabigas, E, Wiesner, J, 2016a. Updated checklist of tiger beetles in the Philippines (Coleoptera, Carabidae, Cicindelidae) (131th. Contribution towards the knowledge of Cicindelidae). Lambillionea 116 (3): 188 - 201" title="Updated checklist of tiger beetles in the Philippines (Coleoptera, Carabidae, Cicindelidae) (131 th. Contribution towards the knowledge of Cicindelidae)." volume="116" year="2016 a">Cabras et al. 2016a</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.11598/btb.2016.23.2.564" author="Cabras, AA" journalOrPublisher="Biotropia" pageId="0" pageNumber="37" pagination="137 - 143" refId="B7" refString="Cabras, AA, Medina, MND, Wiesner, J, 2016b. Tiger beetles (Coleoptera: Carabidae: Cicindelinae) of Compostela Valley Province, Mindanao Island, Philippines. Biotropia 23 (2): 137 - 143, DOI: https://doi.org/10.11598/btb.2016.23.2.564" title="Tiger beetles (Coleoptera: Carabidae: Cicindelinae) of Compostela Valley Province, Mindanao Island, Philippines." url="https://doi.org/10.11598/btb.2016.23.2.564" volume="23" year="2016 b">b</bibRefCitation>
; 
<bibRefCitation author="Cabras, AA" journalOrPublisher="International Research Journal of Biological Sciences" pageId="0" pageNumber="37" pagination="1 - 6" refId="B8" refString="Cabras, AA, Wiesner, J, 2016. Tiger beetles (Coleoptera: Carabidae: Cicindelinae) of Mainit Hotspring and Mati Protected Landscape, Mindanao with notes on their ecology and threats. International Research Journal of Biological Sciences 5 (9): 1 - 6" title="Tiger beetles (Coleoptera: Carabidae: Cicindelinae) of Mainit Hotspring and Mati Protected Landscape, Mindanao with notes on their ecology and threats." volume="5" year="2016">Cabras and Wiesner 2016</bibRefCitation>
; 
<bibRefCitation author="Pepito, J" journalOrPublisher="Journal of Tropical Coleopterology" pageId="0" pageNumber="37" pagination="35 - 46" refId="B47" refString="Pepito, J, Torrejos, C, Cabras, A, Medina, MN, Cudera, R, 2020. Preliminary List of Carabidae and Cicindelidae (Coleoptera) Fauna in Lake Holon, T'boli, South Cotabato, Philippines. Journal of Tropical Coleopterology 1 (2): 35 - 46" title="Preliminary List of Carabidae and Cicindelidae (Coleoptera) Fauna in Lake Holon, T'boli, South Cotabato, Philippines." volume="1" year="2020">Pepito et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="literature data for northern mindanao">
<paragraph pageId="0" pageNumber="37">Literature data for Northern Mindanao.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Bukidnon province</emphasis>
: Impasug-ong (
<bibRefCitation author="Cabras, AA" journalOrPublisher="Lambillionea" pageId="0" pageNumber="37" pagination="188 - 201" refId="B6" refString="Cabras, AA, Cabigas, E, Wiesner, J, 2016a. Updated checklist of tiger beetles in the Philippines (Coleoptera, Carabidae, Cicindelidae) (131th. Contribution towards the knowledge of Cicindelidae). Lambillionea 116 (3): 188 - 201" title="Updated checklist of tiger beetles in the Philippines (Coleoptera, Carabidae, Cicindelidae) (131 th. Contribution towards the knowledge of Cicindelidae)." volume="116" year="2016 a">Cabras et al. 2016a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="materials_examined">
<paragraph pageId="0" pageNumber="37">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Bukidnon province</emphasis>
: Mt. Kitanglad, 28.07.1990, 1♂ 1♀, ex coll. Y. Nishiyama (JWC); Mt. Kitanglad, 11-12.2014, 6♂♂ 6♀♀, leg. N. Mohagan (JWC); Mt. Kalatungan, Sitio Bato, Municipality of Maramag, 11.2019, 1 ex., leg. R. 
<normalizedToken originalValue="Jaskuła">Jaskula</normalizedToken>
(RJC). 
<emphasis bold="true" pageId="0" pageNumber="37">Lanao del Norte province</emphasis>
: Dodiongan Falls, Iligan City - Barangay Bonbonon, 
<geoCoordinate degrees="8.271457" direction="north" orientation="latitude" precision="1" value="8.271457">8.271457N</geoCoordinate>
, 
<geoCoordinate degrees="124.314140" direction="east" orientation="longitude" precision="1" value="124.31414">124.314140E</geoCoordinate>
, 47 m a.s.l., on 
<taxonomicName family="Araceae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="37" rank="family">Araceae</taxonomicName>
leaves, 07.12.2018, 1♂, leg. R. 
<normalizedToken originalValue="Jaskuła">Jaskula</normalizedToken>
(RJC), 11.2019, 6 exx., leg. D. A. P. Acal, J. Ebina, M. L. Lumontod, R. 
<normalizedToken originalValue="Jaskuła">Jaskula</normalizedToken>
(RJC); Mount Agad-agad, 
<geoCoordinate degrees="8" direction="north" minutes="12" orientation="latitude" precision="1" seconds="49.34" value="8.213705">8°12'49.34&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="124" direction="east" minutes="16" orientation="longitude" precision="1" seconds="9.66" value="124.269356">124°16'9.66&quot;E</geoCoordinate>
, ca. 470 m a.s.l., on leaves, 11.2019, 2 exx., D. A. P. Acal, J. Ebina (RJC). 
<emphasis bold="true" pageId="0" pageNumber="37">Misamis Oriental province</emphasis>
: Mimbilisan Protected Landscape, 
<geoCoordinate degrees="8.94884" direction="north" orientation="latitude" precision="1" value="8.94884">8.94884N</geoCoordinate>
, 
<geoCoordinate degrees="124.86517" direction="east" orientation="longitude" precision="1" value="124.86517">124.86517E</geoCoordinate>
, 501 m a.s.l., 18.07.2017, 3 exx., leg. O. Bagona (RJC); Bolyok Falls, Barangay Lubilan, Naawan Municipality, 11.20218, 4 exx., leg. R. 
<normalizedToken originalValue="Jaskuła">Jaskula</normalizedToken>
, D.A.P. Acal, J. Ebina, M. L. Lumontod (RJC); Mambuntan Falls, Barangay Lubilan, Naawan Municipality, 
<geoCoordinate degrees="8.412300" direction="north" orientation="latitude" precision="1" value="8.4123">8.412300N</geoCoordinate>
, 
<geoCoordinate degrees="124.351642" direction="east" orientation="longitude" precision="1" value="124.35164">124.351642E</geoCoordinate>
, 11.2019, 1 ex., leg. R. 
<normalizedToken originalValue="Jaskuła">Jaskula</normalizedToken>
(RJC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="habitat">
<paragraph pageId="0" pageNumber="37">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="37">Forest species noted on tree trunks and leaves.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="remarks">
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="37">
When disturbing, actively fast flying among trees; during flight shows bright orange abdomen coloration. Taxonomical status of both subspecies of 
<taxonomicName authorityName="Erichson" authorityYear="1834" class="Insecta" family="Cicindelidae" genus="Therates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Therates coracinus" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="coracinus">
<emphasis italics="true" pageId="0" pageNumber="37">Therates coracinus</emphasis>
</taxonomicName>
noted in Mindanao (spp. 
<taxonomicName authorityName="Erichson" authorityYear="1834" class="Insecta" family="Cicindelidae" genus="Therates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Therates coracinus" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="coracinus">
<emphasis italics="true" pageId="0" pageNumber="37">Therates coracinus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Wiesner" authorityYear="1988" class="Insecta" family="Cicindelidae" genus="Therates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Therates coracinus subsp. fulvescens" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="coracinus" subSpecies="fulvescens">Therates coracinus ssp. fulvescens</taxonomicName>
Wiesner, 1988) should be revised including molecular data as they probably represent separate species or synonyms.
</paragraph>
</subSubSection>
</treatment>
</document>