Order Rodentia - Family Echimyidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2 1575 1592 book chapter 0-8018-8221-4 10.5281/zenodo.7316535 Proechimys quadruplicatus Hershkovitz 1948 Proechimys quadruplicatus Hershkovitz 1948 , Proc. U. S. Natl. Mus., 97: 138 . Type Locality: Ecuador , Napo-Pastaza Prov., Río Napo , Isla Llunchi (18 kms below mouth of Rio Coca). Vernacular Names: Napo Spiny-rat . Synonyms: Proechimys amphichoricus Moojen 1948 . Distribution: N Peru , E Ecuador and SE Colombia east across S Venezuela and adjacent Brazil to the vicinity of Manaus, west of Rio Negro. Conservation: IUCN – Lower Risk (lc) as P. quadruplicatus and P. amphichoricus . Discussion: Placed in goeldii group by Patton (1987) ; Patton et al. (2000) included amphichoricus . Karyotype has 2n=26-28 and FN=42-44 ( Gardner and Emmons, 1984 ; Patton et al., 2000 ).