The Indo-Paci ® c Pilumnidae XII. On the familial placement of Chlorodiella bidentata (Nobili, 1901) and Tanaocheles stenochilus Kropp, 1984 using adult and larval characters with the establishment of a new subfamily, Tanaochelinae (Crustacea: Decapoda: Brachyura) Author Ng, Peter K. L. text Journal of Natural History 2000 34 207 245 journal article 1464-5262 Pilodius areolatus (H. Milne Edwards, 1834 ) (®gures 12a±e, 13a±c, 14a, b, 15a, b) Material examined. Mauritius : Albion Rocks , Baie de la Petite Rivihre , o Victory Road , Petite Rivihre , Albion , c.20ss12.5¾S 57ss23.5¾E, one m (ovigerous) (spent crab and zoea I, NHM, reg. no. 1998.603), hatched 5 May 1995 , coll. P. Clark by SCUBA , 3±7 m , 6 May 1995 . Zoea I description . Carapace (®gure 12a, d, e): dorsal spine long and curved distally with small tubercules, nearly twice as long as rostral spine; rostral spine much shorter than dorsal spine, approximately equal in length to protopod of antenna, with three spines distally; lateral spines short and straight; one pair of posterodorsal setae; ventral margin without setae; eyes sessile. Antennule (®gure 12b): uniramous, endopod absent; exopod unsegmented with two broad, long, two shorter, slender terminal aesthetascs and one terminal setae. FIG. 11. Phymodius monticulosus : ®rst stage zoea; (a) dorsal view of abdomen; (b) lateral view of abdomen. Antenna (®gure 12a, c): protopodal process distally spinulate, approximatel y equal in length to rostral spine; endopod reduced; exopod rudimentary, unsegmented with 2 unequal terminal setae. Mandible: endopod palp absent. Maxillule (®gure 13a): coxal endite with seven setae; basial endite with ®ve setal processes and two small teeth; endopod two-segmented, proximal segment with one seta; distal segment with six (2 subterminal, 4 terminal) setae; exopod seta absent. Maxilla (®gure 13b): coxal endite bilobed with 4 +4 setae; basial endite bilobed with 5 +4 setae; endopod bilobed, with 3 +5 (2 subterminal and 3 terminal) setae; exopod (scaphognathite) margin with four setae and one long distal stout process. First maxilliped (®gure 14a): coxal segment without setae; basis with 10 setae arranged 2,2,3,3; endopod ®ve-segmented with 3,2,1,2, 5 (1 subterminal and 4 terminal) setae respectively; exopod two-segmented, distal segment with four long terminal plumose natatory setae. FIG. 12. Pilodius areolatus : ®rst stage zoea; (a) anterior view of carapace; (b) antennule; (c) antenna; (d) dorsal carapace spine; (e) rostral spine. Second maxilliped (®gure 14b): coxa without setae; basis with four setae; endopod three-segmented, with 1,1,6 (3 subterminal and 3 terminal) setae respectively; exopod two-segmented, distal segment with 4 long terminal plumose natatory setae. Third maxilliped: absent. Pereiopods: absent. Abdomen (®gure 15a, b): ®ve somites; somite 2 with one pair of dorsolateral processes directed anteriorly; somite 3 with one pair of dorsolateral processes directed ventrally; somites 1±2 with rounded posterolateral processes and 3±5 with short posterolateral spinous processes; somite 1 without setae; somites 2±5 with one pair of posterodorsal setae; pleopod buds absent. FIG. 13. Pilodius areolatus : ®rst stage zoea; (a) maxillule; (b) maxilla; (c) telson. Telson (®gures 13c, 15a, b): each fork long, gradually curved distally; one large and one small lateral spine; one large dorsal medial spine; posterior margin with three pairs of stout spinulate setae.