The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina) Author Erwin, Terry L. text Zootaxa 2002 119 1 68 journal article 10.5281/zenodo.162803 f514e964-2196-461c-84ff-64243643bc81 1175­5326 162803 Agra santarosa Erwin , new species ( Figs. 16a, 16 b, 16c, 16d) Holotype . Male, COSTA RICA : Guanacaste, Santa Rosa National Park, 280m , 10º50'N 85º37'W , May [ 1980 ], (D.H. Janzen & W. Hallwachs) ( USNM : ADP 55010 ). FIGURE 16. A. santarosa , male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0. Diagnosis . Black. Femora rufous, tibiae rufopiceous, knees and tarsi piceous. Elytral interneurs with regularly spaced and slightly cribriform punctulae; apex with more or less similar­sized lateral and sutural dentiform projections. Eyes very large, hemispherical, head short behind eyes. Description . Color and luster: Black. Antennomeres 4­11 bicolored, red and black, tibiae rufous. surface Shiny. Form: Labrum entire. Head ( Fig. 16a ) behind eye moderately short, tapered in both sexes, more so in male. Prothorax "inflated" at basal third. Prothorax subtubular, short, robust, sutures effaced, dorsal surface flattened and densely punctate. Elytral apex ( Fig. 16 b) with more or less similar acute dentiform projections; interneurs with regularly spaced and slightly cribriform punctulae. Sternum VI ( Fig. 16 c) deeply vnotched in male, shallowly v­notched in female. Aedeagus ( Fig. 16 d) with ostium elongate, apex a double scimitar, parameres glabrous. Size: moderately large, 17.5 to 20.0mm in length, 4.86 to 5.16mm in width. Other specimens examined. Costa Rica : 2 paratypes , both females, from the following Guanacaste Conservation Area. Costa Rica , Guanacaste, 1f, June, (D.H. Janzen & W. Hallwachs)( INBIO : ADP55016), 1f, Estacíon Maritza, Volcán de Orosi, East slope, 600m , LN 326900,373000, June, (GNP Biodiversity Survey)( INBIO : CRI 000­134050). Specific epithet. The specific epithet, santarosa , is a noun in apposition based on the type locality, Santa Rosa National Park. Notes. An illustration of famula­group stylomere 2 is on the Agra fortuna web page.