A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae) Author Pimvichai, Piyatida 4D882A89-E0C1-4F78-93E7-FA1B5A2650F2 Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand. & urn: lsid: zoobank. org: author: 4 D 882 A 89 - E 0 C 1 - 4 F 78 - 93 E 7 - FA 1 B 5 A 2650 F 2 & Corresponding author: piyatida _ pimvichai @ yahoo. com piyatida_pimvichai@yahoo.com Author Enghoff, Henrik FB09A817-000D-43C3-BCC4-2BC1E5373635 Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark. & Email: henghoff @ snm. ku. dk & urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635 henghoff@snm.ku.dk Author Panha, Somsak AC935098-D901-4F35-A414-4B0D4FE44E79 Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. & Email: somsakp @ sc. chula. ac. th & urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 somsakp@sc.chula.ac.th Author Backeljau, Thierry E6F42575-36AE-4AD7-98C6-D083EF052568 University of Antwerp, Evolutionary Ecology Group, Groenenborgerlaan 171, B- 2020 Antwerp, Belgium and Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny, and JEMU, Vautierstraat 29, B- 1000 Brussels, Belgium. & Email: Thierry. Backeljau @ naturalsciences. be & urn: lsid: zoobank. org: author: E 6 F 42575 - 36 AE- 4 AD 7 - 98 C 6 - D 083 EF 052568 ackeljau@naturalsciences.be text European Journal of Taxonomy 2016 2016-05-24 199 1 37 journal article 21904 10.5852/ejt.2016.199 d88184f4-ceef-4392-a444-0ac1552778d3 2118-9773 3840484 EE35A675-06C7-4FF9-9EE0-0854D95036C4 Thyropygus planispina sp. nov. urn:lsid:zoobank.org:act: 2F5A25A4-44CC-4B93-AC17-9A9B6E6E9E1F Fig. 7 A–E Diagnosis A species of the opinatus subgroup. Differs from all other species in the subgroup by having the lateral process of the anterior coxal fold ( alp ) regularly curved, terminating in a sharp spine pointing slightly distad, and by having the tibial spine ( ti ) short and flattened. Etymology The name is a Latin noun in apposition and refers to the flattened tibial spine ( ti ). Material examined Holotype THAILAND : ³, Krabi Province , Muang District , Tham Sua temple, 8°07'30" N , 98°55'30" E , 15 Jan. 2012 , leg. P. Pimvichai , C. Sutcharit and members of the Animal Systematics Research Unit ( CUMZ-D00088 ). Fig. 7. Thyropygus planispina sp. nov. , from Tham Sua temple, holotype (CUMZ-D00088), ³, gonopods. A . Anterior view, left telopodite removed. B . Posterior view, left telopodite removed. C . Lateral view. D . Left telopodite, posterior-mesal view. E . Left telopodite, anterior-lateral view. Paratypes THAILAND : 3 ♀♀ , same data as holotype ( CUMZ-D 00097). Description Adult male with 64 podous rings, no apodous rings. Length 13 cm , width 7.6 mm . Adult females with 60–63 podous rings, no apodous rings. Length 13–14 cm , width 7.8–8.6 mm . Colour in life unknown; preserved specimens with head, antennae, legs, metazona, epiproct, paraprocts and hypoproct brown; prozona yellowish brown. GONOPODS ( Fig. 7 A–E). Anterior coxal fold ( ac ; Fig. 7A ): lateral process ( alp ) regularly curved, terminating in sharp spine pointing slightly distad, with lateral short, sharp crest at middle of alp ( Fig. 7C , arrow); mesal process ( amp ) slightly shorter than alp , flattened, straight, directed distad. Posterior coxal fold ( pc ; Fig. 7B ) basally with moderately high lateral paracoxites ( px ), distally truncate, forming shelf for accommodation of telopodite, mesal process ( pmp ) triangular, directed distad, lateral process ( plp ) erect, digitiform. Telopodite ( Fig. 7 D–E) leaving coxite over shelf of posterior coxal fold; femoral spine double, fe 1 very long, curved downward, situated above fe 2 , in situ resting at middle of alp, fe 2 long, very slender, curved downward; tibial spine ( ti ) flattened, short, curving mesad; apical part: spatulate lobe ( sl ) small, rounded; with slender, long spine ( sls ) at base of apical part; palette ( pa ) simple, gutter-like, distally with about nine brownish blepharochaetae ( bp ). DNA barcode The GenBank accession number of the barcode of the holotype is KU306521 (voucher code CUMZ-D00088). Distribution ( Fig. 12 ) Known only from the type locality.