A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae) Author Fery, Hans text Zootaxa 2017 2017-05-05 4317 3 499 529 journal article 32201 10.11646/zootaxa.4317.3.4 ca0d653d-32de-4c75-a95d-d9a4690d6fda 1175-5326 884766 5Fd492A4-D41D-4F37-A121-Ffa680E7E778 Genus Hygrotus Stephens, 1828 Type species : Dytiscus inaequalis Fabricius, 1777 : 239 , by subsequent designation of Curtis (1835: pl. 531) . The genus Hygrotus as here defined includes 129 described species (two of them bitypic) distributed in the Nearctic (including the north of MeXico ), Palaearctic, Afrotropical and Oriental zoogeographical regions ( Fig. 2 and Table 1 ). One Nearctic species was introduced to Hawaii (see below), which belongs to the Pacific region . Diagnosis. In the new classification Hygrotus is the second of two genera of the tribe Hygrotini . As in Clemnius n. gen . all species of the genus have two carinae in each antennal cavity (see Figs 4–6 ). According to the two main clades in the phylogeny of Hygrotini and several morphological characters, the genus (which corresponds to Clade B in Fig. 1 ) is subdivided into four clades corresponding to the following four subgenera: clade B1 = subgenus Coelambus , clade B2 = subgenus Hyphoporus , clade B3 = subgenus Hygrotus s. str. and clade B4 = Leptolambus n. subgen.