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<mods:title id="A0BF91D797FD6EA8A01794BAF4264EE1">Rediscovery and re (re) description of Scydmaenus kasuganus Franz, the only Japanese species of the subgenus Mascarensia Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<taxonomicName id="4C12ACF5FFABFF87FF5FF982C4C1F998" ID-CoL="4W3W4" authority="Franz" authorityName="Franz" authorityYear="1975" box="[151,727,1626,1653]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="australiensis" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF5FF982C739F998" bold="true" box="[151,303,1626,1653]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE88F983C7CDF998" bold="true" box="[320,475,1627,1653]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE24F982C493F998" bold="true" box="[492,645,1626,1653]" italics="true" pageId="7" pageNumber="45">australiensis</emphasis>
Franz
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[
<collectingCountry id="F30597E6FFABFF87FD2EF982C577F999" box="[742,865,1626,1652]" name="Australia" pageId="7" pageNumber="45">Australia</collectingCountry>
]
</emphasis>
</heading>
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<emphasis id="B9660B64FFABFF87FF0FF9A3C752F97E" box="[199,324,1659,1683]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE9BF9A4C7C3F97E" box="[339,469,1660,1683]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE2CF9A3C470F97E" box="[484,614,1659,1683]" italics="true" pageId="7" pageNumber="45">australiensis</emphasis>
<bibRefCitation id="EF83AA87FFABFF87FDA4F9A4C53EF97E" author="Franz, H." box="[620,808,1659,1683]" pageId="7" pageNumber="45" pagination="1 - 312" refId="ref5783" refString="Franz, H. (1975 b) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975b: 282</bibRefCitation>
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.
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<emphasis id="B9660B64FFABFF87FF5FF945C554F95A" bold="true" box="[151,834,1693,1720]" pageId="7" pageNumber="45">
<taxonomicName id="4C12ACF5FFABFF87FF5FF945C4C8F95A" ID-CoL="4W3W5" authority="Franz" authorityName="Franz" authorityYear="1973" box="[151,734,1693,1720]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dendrophilus" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF5FF945C739F955" bold="true" box="[151,303,1693,1720]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE88F945C7CDF95A" bold="true" box="[320,475,1693,1719]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE24F945C49AF955" bold="true" box="[492,652,1693,1720]" italics="true" pageId="7" pageNumber="45">dendrophilus</emphasis>
Franz
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[
<collectingCountry id="F30597E6FFABFF87FD25F945C52DF95A" box="[749,827,1693,1719]" name="Nepal" pageId="7" pageNumber="45">Nepal</collectingCountry>
]
</emphasis>
</paragraph>
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<emphasis id="B9660B64FFABFF87FF0FF919C747F936" box="[199,337,1729,1755]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FEAAF91AC7E4F936" box="[354,498,1730,1755]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FDCAF919C483F936" box="[514,661,1729,1755]" italics="true" pageId="7" pageNumber="45">dendrophilus</emphasis>
<bibRefCitation id="EF83AA87FFABFF87FD54F91AC54BF936" author="Franz, H." box="[668,861,1729,1756]" pageId="7" pageNumber="45" pagination="113 - 156" refId="ref5702" refString="Franz, H. (1973) Die auf meinen Forschungsreisen nach Nepal in den Jahren 1970 und 1971 gesammelten Scydmaeniden und einige nordindische Vertreter dieser Familie (Coleoptera, Scydmaenidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 23, 113 - 156. [1971]" type="journal article" year="1973">Franz, 1973: 120</bibRefCitation>
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<emphasis id="B9660B64FFABFF87FF5FF93DC5B2F912" bold="true" box="[151,932,1765,1792]" pageId="7" pageNumber="45">
<taxonomicName id="4C12ACF5FFABFF87FF5FF93DC4B9F912" ID-CoL="4W3W6" authority="Franz" authorityName="Franz" authorityYear="1992" box="[151,687,1765,1792]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dissimilis" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF5FF93DC739F8ED" bold="true" box="[151,303,1765,1792]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE88F93DC7CDF912" bold="true" box="[320,475,1765,1791]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE24F93DC44AF8ED" bold="true" box="[492,604,1765,1792]" italics="true" pageId="7" pageNumber="45">dissimilis</emphasis>
Franz
</taxonomicName>
[
<collectingCountry id="F30597E6FFABFF87FD75F93DC524F912" box="[701,818,1765,1791]" name="Malaysia" pageId="7" pageNumber="45">Malaysia</collectingCountry>
(
<collectingRegion id="49D61994FFABFF87FC89F93DC582F912" box="[833,916,1765,1791]" country="Malaysia" name="Sabah" pageId="7" pageNumber="45">Sabah</collectingRegion>
)]
</emphasis>
</paragraph>
<paragraph id="8BADD776FFABFF87FF0FF8DEC4EBF8F3" blockId="7.[151,1436,1626,2017]" box="[199,765,1798,1822]" pageId="7" pageNumber="45">
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<treatmentCitation id="0AB3F167FFABFF87FF0FF8DEC4EFF8F3" author="Franz, H." box="[199,761,1798,1822]" page="874" pageId="7" pageNumber="45" year="1992">
<taxonomicName id="4C12ACF5FFABFF87FF0FF8DEC4EFF8F3" ID-CoL="4W3W6" authority="Franz, 1992: 874" authorityName="Franz" authorityPageNumber="874" authorityYear="1992" box="[199,761,1798,1822]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dissimilis" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF0FF8DEC752F8F3" box="[199,324,1798,1822]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE9BF8DEC7C3F8F0" box="[339,469,1798,1821]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE2CF8DEC452F8F3" box="[484,580,1798,1822]" italics="true" pageId="7" pageNumber="45">dissimilis</emphasis>
<bibRefCitation id="EF83AA87FFABFF87FD82F8DEC4EFF8F3" author="Franz, H." box="[586,761,1798,1822]" pageId="7" pageNumber="45" pagination="859 - 953" refId="ref6002" refString="Franz, H. (1992) Monographie der Scydmaenidae (Coleoptera) von Sabah (NO-Borneo). Revue suisse de Zoologie, 99 (4), 859 - 953. https: // doi. org / 10.5962 / bhl. part. 79857" type="journal article" year="1992">Franz, 1992: 874</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BADD776FFABFF87FF5FF8FFC5B2F8AF" blockId="7.[151,1436,1626,2017]" box="[151,932,1831,1858]" pageId="7" pageNumber="45">
<emphasis id="B9660B64FFABFF87FF5FF8FFC5B2F8AF" bold="true" box="[151,932,1831,1858]" pageId="7" pageNumber="45">
<taxonomicName id="4C12ACF5FFABFF87FF5FF8FFC4D7F8AF" ID-CoL="4W3W8" authority="Franz" authorityName="Franz" box="[151,705,1831,1858]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="kasuganus" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF5FF8FFC739F8AF" bold="true" box="[151,303,1831,1858]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE88F8F0C7CDF8AF" bold="true" box="[320,475,1832,1858]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE24F8FFC478F8AF" bold="true" box="[492,622,1831,1858]" italics="true" pageId="7" pageNumber="45">kasuganus</emphasis>
Franz
</taxonomicName>
[
<collectingCountry id="F30597E6FFABFF87FD07F8F0C508F8AF" box="[719,798,1832,1858]" name="Japan" pageId="7" pageNumber="45">Japan</collectingCountry>
(Honshu)]
</emphasis>
</paragraph>
<paragraph id="8BADD776FFABFF87FF0FF890C777F870" blockId="7.[151,1436,1626,2017]" pageId="7" pageNumber="45">
<treatmentCitationGroup id="AB02F058FFABFF87FF0FF890C777F870" pageId="7" pageNumber="45">
<treatmentCitation id="0AB3F167FFABFF87FF0FF890C4EEF88D" author="Franz, H." box="[199,760,1864,1889]" page="58" pageId="7" pageNumber="45" year="1976">
<taxonomicName id="4C12ACF5FFABFF87FF0FF890C4EEF88D" ID-CoL="4W3W8" authority="Franz, 1976: 58" authorityName="Franz" authorityPageNumber="58" authorityYear="1976" box="[199,760,1864,1889]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="kasuganus" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF0FF890C752F88D" box="[199,324,1864,1888]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE9BF891C7C3F88D" box="[339,469,1865,1888]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE2CF890C459F88D" box="[484,591,1864,1888]" italics="true" pageId="7" pageNumber="45">kasuganus</emphasis>
<bibRefCitation id="EF83AA87FFABFF87FD9DF891C4EEF88D" author="Franz, H." box="[597,760,1864,1888]" pageId="7" pageNumber="45" pagination="51 - 60" refId="ref5837" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Ergebnisse einer auf Einladung der Internationalen Vereinigung fur Vegetationskunde nach Japan durchgefuhrten Forschungsreise. Entomologische Blatter fur Biologie und Systematik der Kafer, 72 (1), 51 - 60." type="journal article" year="1976">Franz, 1976: 58</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Redescribed by 
<treatmentCitation id="0AB3F167FFABFF87FC6DF890C242F88C" author="Hoshina, H." box="[933,1108,1864,1889]" page="2" pageId="7" pageNumber="45" year="2007">
<bibRefCitation id="EF83AA87FFABFF87FC6DF890C242F88C" author="Hoshina, H." box="[933,1108,1864,1889]" pageId="7" pageNumber="45" pagination="1 - 6" refId="ref6049" refString="Hoshina, H. (2007) Redescriptions of three species of the genus Scydmaenus (Coleoptera: Scydmaenidae) from Japan and Taiwan. Memoirs of the Faculty of Education and Regional Studies, Fukui University, 2: Natural Science, 58, 1 - 6." type="journal article" year="2007">Hoshina, 2007: 2</bibRefCitation>
</treatmentCitation>
, and in present paper. 
<treatmentCitation id="0AB3F167FFABFF87FF0FF8BFC51BF892" author="Franz, H." box="[199,781,1895,1919]" page="57" pageId="7" pageNumber="45" year="1976">
<taxonomicName id="4C12ACF5FFABFF87FF0FF8BFC51BF892" authority="Franz, 1976: 57" authorityName="Franz" authorityPageNumber="57" authorityYear="1976" box="[199,781,1895,1919]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="honshuensis" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF0FF8BFC752F892" box="[199,324,1895,1919]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE9CF8B0C7C0F892" box="[340,470,1896,1919]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE2EF8BFC474F892" box="[486,610,1895,1919]" italics="true" pageId="7" pageNumber="45">honshuensis</emphasis>
<bibRefCitation id="EF83AA87FFABFF87FDA1F8B0C51BF892" author="Franz, H." box="[617,781,1895,1919]" pageId="7" pageNumber="45" pagination="51 - 60" refId="ref5837" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Ergebnisse einer auf Einladung der Internationalen Vereinigung fur Vegetationskunde nach Japan durchgefuhrten Forschungsreise. Entomologische Blatter fur Biologie und Systematik der Kafer, 72 (1), 51 - 60." type="journal article" year="1976">Franz, 1976: 57</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Apparent lapsus for 
<emphasis id="B9660B64FFABFF87FC20F8BFC245F892" box="[1000,1107,1895,1919]" italics="true" pageId="7" pageNumber="45">kasuganus</emphasis>
; name not mentioned anywhere else in the text.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BADD776FFABFF87FF5FF87FC224F82F" blockId="7.[151,1436,1626,2017]" box="[151,1074,1959,1986]" pageId="7" pageNumber="45">
<emphasis id="B9660B64FFABFF87FF5FF87FC224F82F" bold="true" box="[151,1074,1959,1986]" pageId="7" pageNumber="45">
<taxonomicName id="4C12ACF5FFABFF87FF5FF87FC497F82F" ID-CoL="4W3W9" authority="Franz" authorityName="Franz" authorityYear="1982" box="[151,641,1959,1986]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="loebli" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF5FF87FC739F82F" bold="true" box="[151,303,1959,1986]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE88F870C7CDF82F" bold="true" box="[320,475,1960,1986]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE24F87FC439F82F" bold="true" box="[492,559,1959,1986]" italics="true" pageId="7" pageNumber="45">loebli</emphasis>
Franz
</taxonomicName>
[
<collectingCountry id="F30597E6FFABFF87FD47F870C51AF82F" box="[655,780,1960,1986]" name="Indonesia" pageId="7" pageNumber="45">Indonesia</collectingCountry>
(
<collectingRegion id="49D61994FFABFF87FCD4F870C598F82F" box="[796,910,1960,1986]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="45">Sumatra</collectingRegion>
), 
<collectingCountry id="F30597E6FFABFF87FC6BF870C230F82F" box="[931,1062,1960,1986]" name="Sri Lanka" pageId="7" pageNumber="45">Sri Lanka</collectingCountry>
].
</emphasis>
</paragraph>
<paragraph id="8BADD776FFABFF87FF0FF810C4C1F80C" blockId="7.[151,1436,1626,2017]" box="[199,727,1992,2017]" pageId="7" pageNumber="45">
<treatmentCitationGroup id="AB02F058FFABFF87FF0FF810C4C1F80C" box="[199,727,1992,2017]" pageId="7" pageNumber="45">
<treatmentCitation id="0AB3F167FFABFF87FF0FF810C4C5F80C" author="Franz, H." box="[199,723,1992,2017]" page="133" pageId="7" pageNumber="45" year="1982">
<taxonomicName id="4C12ACF5FFABFF87FF0FF810C4C5F80C" ID-CoL="4W3W9" authority="Franz, 1982: 133" authorityName="Franz" authorityPageNumber="133" authorityYear="1982" box="[199,723,1992,2017]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="loebli" subGenus="Mascarensia">
<emphasis id="B9660B64FFABFF87FF0FF810C752F80D" box="[199,324,1992,2016]" italics="true" pageId="7" pageNumber="45">Scydmaenus</emphasis>
(
<emphasis id="B9660B64FFABFF87FE9BF811C7C3F80D" box="[339,469,1993,2016]" italics="true" pageId="7" pageNumber="45">Mascarensia</emphasis>
) 
<emphasis id="B9660B64FFABFF87FE2CF810C408F80D" box="[484,542,1992,2016]" italics="true" pageId="7" pageNumber="45">loebli</emphasis>
<bibRefCitation id="EF83AA87FFABFF87FDECF811C4C5F80C" author="Franz, H." box="[548,723,1993,2017]" pageId="7" pageNumber="45" pagination="125 - 274" refId="ref5888" refString="Franz, H. (1982) Coleoptera: Die Scydmaenidae Sri Lankas (mit Ausnahme der Genera Cephennium s. lat., Clidicus und Syndicus). Entomologica Scandinavica, Supplement 11, 125 - 274." type="journal article" year="1982">Franz, 1982: 133</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>