Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae) Author De Mattia, Willy https://orcid.org/0000-0002-0056-467X Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria willy.demattia@icgeb.org Author Reier, Susanne Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria Author Haring, Elisabeth Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria text ZooKeys 2021 2021-12-14 1077 1 175 http://dx.doi.org/10.3897/zookeys.1077.67081 journal article http://dx.doi.org/10.3897/zookeys.1077.67081 1313-2970-1077-1 C28AD65A76F242CFBED7DFB3702CABCE 734088641608531C8E2CC69397B000ED Charpentieria itala baldensis (Strobel, 1851) Figs 45.1-45.3, 47.7 Distribution. Charpentieria itala baldensis is exclusively known from Monte Baldo and Monte Altissimo (Verona, Veneto, Italy), restricted to the peak areas. Figure 44. Charpentieria itala itala (G. von Martens, 1824), Barbarano Vicentino 44.1, 44.2 whole distal genital organs 44.3 internal distal part of genital organs 44.4 detail of the pseudopapilla. Charpentieria itala albopustolata (De Cristofori & Jan, 1832), Brescia 44.5 whole distal genital organs 44.6 internal distal part of genital organs. Charpentieria itala allatollae ( Kaeufel , 1928), Val Ampola, Storo 44.7 whole distal genital organs 44.8 internal distal part of genital organs. Specimens examined. Italy , Veneto , Verona , Naole , San Zeno di Montagna. 900 m asl , 45°44'25.95"N , 10°51'13.69"E , [ Lab ID C2_2, COI: MW758955 ], I. Niero leg. and det., 2 dissected spm . External morphology of the genital organs (Fig. 45 .1). The FO is as long as the V. The FDBC is longer than the BC+SDBC (SDBC+BC). The BC+SDBC presents a very distinguishable second duct as the BC abruptly inflates. The apex is rounded. The D is slightly longer than the BC+SDBC, with a thin proximal part and a swollen distal part. The V is long and thin in diameter. The PC is double as long as the V. The P is slightly swollen at its proximal part. The transition area P-E is clearly detectable from outside. The PR is short and robust. The E is slightly longer than the P but thinner in diameter. Internal morphology of the genital organs (Figs 45 .2, 45.3). The A and the V show smooth few, very loose and barely visible longitudinal elevations. The P shows five to six smooth longitudinal plates that proximally directly start from the ER. The PP is smooth, very elongated with a pointed apex and it occupies almost the whole penial internal volume. Its basis is slightly swollen. The PP originates independently, not connecting neither with the ER or the ELP. The epiphallar formula is: ER+PP+ELP. The E shows two remarkably fringed ELP. They proximally fade before the VD. Figure 45. Charpentieria itala baldensis (Strobel, 1851), San Zeno, Verona 45.1 whole distal genital organs 45.2 internal distal part of genital organs 45.3 cross section of the pseudopapilla. Charpentieria itala balsamoi (Strobel, 1850), Val Serina, Bracca 45.4 whole distal genital organs 45.5 internal distal part of genital organs. Remarks. This small subspecies, found exclusively at higher altitudes, was previously considered as a transitional form between C. itala albopustolata and C. itala rubiginea ( Nordsieck 1963b : 176-177). The same author ( Nordsieck 2007 : 54) recently elevated it (with no taxonomic explanation) as a valid subspecies. The taxon was described by Strobel (1851 : 25) and not by Charpentier (1852: 384) as stated in MolluscaBase (2021).