Manual of North American Agromyzidae (Diptera, Schizophora), with revision of the fauna of the " Delmarva " states
Author
Lonsdale, Owen
Agriculture & Agri-Food Canada, 960 Carling Avenue, Ottawa, ON, K 1 A 0 C 6, Canada
neoxabea@hotmail.com
text
ZooKeys
2021
2021-07-29
1051
1
481
http://dx.doi.org/10.3897/zookeys.1051.64603
journal article
http://dx.doi.org/10.3897/zookeys.1051.64603
1313-2970-1051-1
639E252D43924ABB910BCEA5D8AD2487
BE8CC6847F645F61BB2F7A6BCF96FD64
Phytomyza
Fallen
Phytomyza
Fallen
, 1810: 10. Type species:
Phytomyza flaveola
Fallen
, 1810 [=
Musca ranunculi
Schrank, 1803], by monotypy.
Frick 1952a
: 421,
1959
: 420;
Spencer 1969
: 218;
Spencer and Steyskal 1986b
: 172;
Winkler et al. 2009
: 260;
Papp and
Cerny
2020
: 175.
Phytomyia
. Misspelling. Haliday, 1833: 150.
Napomyza
Curtis, 1837: 282 [attributed to Haliday manuscript name]. Type species:
Phytomyza nigricornis
Macquart 1835
, by monotypy.
Napomyza
Westwood, 1840: 152 [as subgenus of
Phytomyza
]. Type species:
Phytomyza festiva
Meigen, 1830 [=
Phytomyza elegans
Meigen, 1830], by monotypy by first reviser action of
Hendel (1920)
.
Hendel 1920
: 111 [attrib. to Haliday; as genus];
Frick 1952a
: 419,
1959
: 419;
Spencer 1969
: 210;
Griffiths and Steyskal 1986
: 170 [proposed supression
Napomyza
Curtis];
Spencer and Steyskal 1986b
: 167;
Zlobin 1994
: 289; ICZN 1988: 77 [suppression of
Napomyza
Curtis];
Winkler et al. 2009
: 271 [as subgenus];
Papp and
Cerny
2020
: 121.
Chromatomyia
Hardy, 1849: 390. Type species
Phytomyza periclymeni
de Meijere, 1924 (misidentified as
Phytomyza obscurella
Fallen
, 1823 by Hardy and Coquillett), by subsequent designation (
Coquillett 1910
: 523) - see discussion in
Griffiths (1974
: 36). This name is not preoccupied by
Chromatomyia
Walker, as
Walker's
name is not available.
Braschnikov 1897
: 40;
Coquillett 1910
: 523;
Frick 1952a
: 421 [as synonym of
Phytomyza
];
Griffiths 1974
: 36;
Spencer and Steyskal 1986b
: 173;
Spencer 1987
: 255;
Papp 1984
: 315 [as synonym of
Phytomyza
];
Spencer and Martinez 1987
: 255 [stat reinst.];
Winkler et al. 2009
: 276 [as synonym of
Phytomyza
];
Papp and
Cerny
2020
: 19;
von Tschirnhaus 2021
: 105.
Dineura
Lioy, 1864: 1315. Type species:
Phytomyza festiva
Meigen, 1830 [=
Phytomyza elegans
Meigen 1830
], by original designation. Preoccupied by Dahlbom (1835) and Selys (1859).
Napomyia
. Misspelling. Schiner, 1868: 227.
Phythomyza
. Misspelling. Rondani, 1874: 51.
Lonicera
. Error for
Phytomyza
.
Meijere 1924
: 147 [in describing and naming
Phytomyza luteoscutellata
Meijere on page 147, the genus group name of the host (
Lonicera
) was used for the fly genus; the correct combination is given on pages 126 and 147].
Ptochomyza
Hering, 1942: 530. Type species:
Ptochomyza asparagi
Hering 1942, by original designation.
Winkler et al. 2009
: 282 [as subgenus];
Lonsdale 2015
: 637.
Phytomyza
is the most speciose genus of
Agromyzidae
with ca. 800 species worldwide, and certainly the most diverse with respect to morphology and host use. The classification of the group was developed by Winker et al. (2009) when they synonymized
Chromatomyia
, and included
Napomyza
and the small genus
Ptochomyza
as subgenera, finding that all of these rendered the larger
Phytomyza
paraphyletic. Further discussion is provided in
Lonsdale (2015)
and
Lonsdale and Eiseman (2021)
.
Species of the
subgenus Napomyza
occur globally, occurring mostly towards the west in North America, and while none are yet known from the Delmarva states, it is likely that they will eventually be found.
Zlobin (1994)
discussed the characters used by Spencer to diagnose
Napomyza
, finding them to be plesiomorphic and not sufficient to maintain the genus as a separate entity. Zlobin instead provided tentative characters of the male genitalia, although similar characters are found in some
Phytomyza
s. s. and should be used with caution (
Winkler et al. 2009
). Species of the
subgenus Ptochomyza
are mostly restricted to the Palaearctic Region; a minority also occur in the Afrotropics, as discussed by
Lonsdale (2015)
.
In previous keys and diagnoses, the length of the fronto-orbital setae and the lengths of the costal sectors were used to differentiate those species of
Phytomyza
with a dark frons. These characters appear to vary much more widely than previously appreciated among many of those species, however, making them unreliable for diagnosis and they are not used in the key provided below. Male dissections should be made whenever possible to verify identifications, as genitalic characters in this genus are among the most complex and distinct of any
Agromyzidae
.