Disentangling the diversity and taxonomy of Hymenophyllaceae (Hymenophyllales, Polypodiidae) in the Comoros
Author
Ahamada H. Saïd
Author
Sabine Hennequin
Author
Germinal Rouhan
Author
Jean-Yves Dubuisson
text
European Journal of Taxonomy
2017
2017-04-28
313
1
53
journal article
32053
10.5852/ejt.2017.313
e9b8382f-faec-4925-993b-f13e4cfa2a71
2118-9773
896401
Crepidomanes
inopinatum
(Pic.Serm.) J.P.Roux
Crepidomanes
inopinatum
(Pic.Serm.) J.P.Roux,
Conspectus
of southern African pteridophyta
: 45 (2001). –
Vandenboschia
inopinata
Pic.Serm.
,
Bulletin du Jardin botanique national de Belgique
53, 1/2: 245, 246 (1983).– Type: DRC, s. loc., s.d.,
R.E.G. Pichi Sermolli 4455
(holo-: FI-PS 19455; iso-: K000435648!, BR 0000008054661!).
Trichomanes
inopinatum
(Pic.Serm.) J.E.Burrows
,
Southern African ferns and fern allies
: 93 (1990).
Trichomanes pyxidiferum
L.f.
majus
Taton
,
Bulletin de la Société royale de Botanique de Belgique
78: 34 (1946). – Type: DRC, s. loc., s.d.,
J.C.C. Bequaert 3931bis
(syn-: BR 0000013208462!); DRC, s. loc., s.d.,
J.C.C. Bequaert 4231
(syn-: BR 0000013204853!); DRC, s. loc., s.d.,
J.C.C. Bequaert s.n
. (syn-: BR 0000013204877!); DRC, s. loc., s.d.,
J.C.C. Bequaert s.n
. (syn-: BR 0000013204860!).
Short description
Epiphytic or lithophytic; long-creeping filiform rhizomes, rootless and densely covered by dense black trichomes, bearing widely separated pending fronds; fronds well stipitate, with wing on the upper part of the stipe, up to 10(–15) cm long, lanceolate to ovate, pinnate-pinnatifid to bipinnate-pinnatifid with sometimes, when dry, folds parallel to veins; sori numerous, paratactic, mostly as long as wide, obconic to campanulate with a slightly dilated mouth.
Global distribution
Central Africa to the western Indian Ocean.
Distribution in Comoros
Grande Comore.
Additional specimens examined
COMOROS: GRANDE COMORE, Massif de la Grille, 30 Nov. 2002,
F.Rakotondrainibe et al. 6841
,
6841bis
(P00312119, P00312120).
Note
The medium-sized (with bipinnate-pinnatifid fronds usually up to 10 cm long,rarely longer)
C. inopinatum
is usually found in highland rainforests from Central Africa to the western Indian Ocean (
Kornaś 1994
) and is here newly reported from the Comoros. It is often confused with the morphologically closely related
C.
melanotrichum
(Schltdl.) J.P.Roux
which differs only by the dense and black indumentum all along the stipe (vs restricted to the stipe base in
C. inopinatum
). Such confusion explains that the BR isotype specimen (BR 0000008054661) was identified on the sheet as
C. melanotrichum
by Beentje in 2007. The two existing Comorian collects do not exhibit the particular dense indumentum all along the stipe suggesting that they should rather represent
C. inopinatum
. But as discussed by
Dubuisson
et al.
(2013b)
, the distinction between
C. inopinatum
and
C. melanotrichum
based on stipe indumentum is not always obvious and thus questionable.