Review of the Chinese species of Eupoecilia Stephens (Lepidoptera: Tortricidae: Cochylini) Author Zhang, Xu Author Li, Houhun text Zootaxa 2008 1692 55 68 journal article 10.5281/zenodo.180634 73a052dc-f882-4b4a-a077-ecced32422b8 1175-5326 180634 Key to Eupoecilia species of the Oriental and Palaearctic regions 1 Male ............................................................................................................................................................ 2 – Female ....................................................................................................................................................... 28 2 Median process of transtilla broader than width of socius; socius well sclerotized ................................... 3 – Median process of transtilla narrower than or equal to width of socius; socius weakly sclerotized ........ 21 3 Aedeagus with comblike distal lobes projecting dorsally .......................................................................... 4 – Aedeagus without such comblike lobes ...................................................................................................... 6 4 Socius distinctly dentate at base ( Razowski, 1968: fig. 22 ) ...................................... E. dentana Razowski – Socius not dentate at base ........................................................................................................................... 5 5 Vesica with one large cornutus almost equal in length to aedeagus ( Razowski, 1968: fig. 25 ) .................... ............................................................................................................................ E. amphimnesta (Meyrick) – Vesica with one large cornutus about half the length of aedeagus ( Razowski, 1968: fig. 21 ) ...................... ................................................................................................................................... E. armifera Razowski 6 Sacculus with spines ................................................................................................................................... 7 – Sacculus without spines ............................................................................................................................ 12 7 Spines on sacculus thick, about 1/2 width of socius ................................................................................... 8 – Spines on sacculus slender, about 1/5 width of socius ............................................................................... 9 8 Vesica with a large cornutus at the base and one at the middle; costa of valva gently concave ( Diakonoff, 1984: figs. 13, 14 ) ...................................................................................................... E. coniopa Diakonoff – Vesica without large cornutus at base and middle; costa of valva nearly straight ( Diakonoff, 1983: figs. 5, 6 ) ............................................................................................................................... E. anebrica Diakonoff 9 Aedeagus with distal cornuti having flat base .......................................................................................... 10 – Aedeagus with distal cornuti lacking flat base (fig. 2b) ........................................ E. ambiguella (Hübner) 10 Vesica with three strong cornuti near middle; central part of transtilla narrow, 2.0-2.5 times width of socius ........................................................................................................................................................ 11 – Vesica with group of more than ten slender cornuti near middle; central part of transtilla broad, 3.5 times width of socius ( Razowski, 1968: figs. 4, 5 ) ................................................... E. tenggerensis (Diakonoff) 11 Cornuti having flat base; central part of transtilla twice width of socius ( Razowski, 1968: figs. 7, 8 ) ......... ................................................................................................................................ E. sumbana (Diakonoff) – Cornuti lacking flat base; central part of transtilla 2.5 times width of socius ( Razowski, 1968: figs. 10, 11 ) .......................................................................................................................... “ E. reliquatrix (Meyrick) ”. 12 Valva broadly subquadrate ........................................................................................................................ 13 – Valva subtriangular or oblong ................................................................................................................... 14 13 Vesica with the largest cornutus about 4/5 of length of aedeagus ( Diakonoff, 1983: fig. 4 ) ......................... ................................................................................................................................ E. sumatrana Diakonoff – Vesica with the largest cornutus about 1/6 of length of aedeagus (fig. 3b) ..................... E. inouei Kawabe 14 Valva subtriangular ................................................................................................................................... 15 – Valva oblong ............................................................................................................................................. 20 15 Vesica with distal cornuti .......................................................................................................................... 16 – Vesica without distal cornuti ..................................................................................................................... 19 16 Vesica with groups of small cornuti at base and near middle ( Diakonoff, 1984: fig. 10 ) ............................. ............................................................................................................................... E. engelinae (Diakonoff) – Vesica with one large cornutus at base or middle ..................................................................................... 17 17 Vesica with cornutus at base ..................................................................................................................... 18 – Vesica with cornutus at middle ( Razowski, 2005: fig. 4 ) ....................................... E. yubariana Razowski 18 Aedeagus twice length of valva, with an apical projection (fig. 3d). E. sanguisorbana (Herrich-Schäffer) – Aedeagus 1.25 times length of valva, without apical projection (fig. 2c) ............... E. angustana (Hübner) 19 Vesica with three strong cornuti; central part of transtilla narrow, twice width of socius ( Razowski, 2005: figs. 5, 6 ) ........................................................................................................................... E. lata Razowski – Vesica with one strong cornutus; central part of transtilla broad, 3.5 times width of socius ( Razowski, 1968: figs. 16, 17 ) ....................................................................................................... E. crocina Razowski 20 Vesica with one straight cornutus extending from base to near middle, distal cornuti capitately arranged (fig. 3a) ..................................................................................................................... E. citrinana Razowski – Vesica with one bent cornutus extending from base to apex, distal cornuti absent ( Razowski, 1970a: Tafel 76: fig. 171 ) ................................................................................................................ E. cebrana (Hübner) 21 Central part of transtilla narrower than width of socius ........................................................................... 22 – Central part of transtilla about equal to width of socius ........................................................................... 24 22 Cucullus as wide as base of valva ( Razowski, 1968: fig. 41 ) .............................. E. charixantha (Meyrick) – Cucullus narrower than base of valva ....................................................................................................... 23 23 Vesica with one large cornutus (fig. 3c) .................................................................... E. kobeana Razowski – Vesica lacking large cornutus ( Razowski, 1968: fig. 45 ) ....................................... E. eucalypta (Meyrick) 24 Sacculus bearing prominence with denticulate ventral edge .................................................................... 25 – Sacculus lacking such prominence ........................................................................................................... 26 25 Vesica with four strong distal cornuti, with a subapical and a second spiny projection; costa of valva nearly straight ( Diakonoff, 1984: figs. 11, 12 ) ....................................................... E. wegneri (Diakonoff) – Vesica with five strong distal cornuti, with one subapical projection only; costa of valva gently concave (fig. 2a) ..................................................................................................................... E. quinaspinalis sp. n. 26 Sacculus with small spines posteriorly; aedeagus broadly parallel-sided, vesica with a cornutus about 2/3 length of aedeagus ( Razowski, 1968: figs. 35, 36 ) ................................................. E. ochrotona Razowski – Sacculus without spines; aedeagus elongate pear-shaped, vesica with a cornutus about half length of aede- agus ............................................................................................................................................................ 27 27 Cucullus oblong ( Diakonoff, 1982: fig. 1 ) ........................................................... E. anisoneura Diakonoff – Cucullus subtriangular ( Diakonoff, 1982: pl. I, fig. 1 ) .............................................. E. cracens Diakonoff 28 Ductus bursae broadened posteriorly ........................................................................................................ 29 – Ductus bursae not broadened posteriorly .................................................................................................. 32 29 Corpus bursae entirely spined ( Diakonoff, 1984: fig. 15 ) ......................................... E. coniopa Diakonoff – Corpus bursae partially spined .................................................................................................................. 30 30 Corpus bursae with long spines forming a “star” at middle ..................................................................... 31 – Corpus bursae without such spines at middle (fig. 4b) .......................................... E. ambiguella (Hübner) 31 Ductus bursae broad, about 2/3 width of corpus bursae; corpus bursae with row of long spines laterally ( Razowski, 1968: fig. 6 ) .................................................................................. E. tenggerensis (Diakonoff) – Ductus bursae narrow, about 1/3 width of corpus bursae; corpus bursae with spines irregularly scattered ( Razowski, 1968: fig. 12 ) ...................................................................................... E. reliquatrix (Meyrick) 32 Ductus bursae length less than twice its greatest width ............................................................................ 33 – Ductus bursae length greater than twice its greatest width ....................................................................... 40 33 Sterigma with denticulate edge posteriorly .............................................................................................. 34 – Sterigma without denticulate edge ............................................................................................................ 36 34 Ductus bursae narrow, about 1/2 width of corpus bursae; corpus bursae with spines forming radiating arc at the anterior edge .................................................................................................................................... 35 – Ductus bursae broad, about 3/4 width of corpus bursae; corpus bursae with spines not forming radiating arc at the anterior edge ( Razowski, 1968: fig. 27 ) .................................................. E. wegneri (Diakonoff) 35 Ductus seminalis originating from anterior 1/3 of ductus bursae (fig. 4a) ............... E. quinaspinalis sp. n. – Ductus seminalis originating from proximal anterior portion of ductus bursae ( Diakonoff, 1984: fig. 16 ). ....................................................................................................................................... E. thalia Diakonoff 36 Ductus bursae entirely membranous ......................................................................................................... 37 – Ductus bursae partially sclerotized ........................................................................................................... 38 37 Sterigma protruding at middle on anterior margin; ductus bursae 1/2 length of corpus bursae ( Razowski, 1968: fig. 26 ) ..................................................................................................... E. amphimnesta (Meyrick) – Sterigma concave at middle on posterior margin; ductus bursae 1/3 length of corpus bursae ( Diakonoff, 1971: fig. 15 ) ..................................................................................................... E. dynodesma (Diakonoff) 38 Apophyses about 1/3 width of papilla analis ( Razowski, 1970a: Tafel 141: fig. 171 ) E. cebrana (Hübner) – Apophyses about 1/6 width of papilla analis ............................................................................................ 39 39 Ductus bursae about 2/3 sclerotized laterally and posteriorly ( Razowski, 1970a: Tafel 141: fig. 172 ) ........ .................................................................................................................................. E. angustana (Hübner) – Ductus bursae about 1/2 sclerotized anteriorly ( Razowski, 1968: fig. 9 ) .............. E. sumbana (Diakonoff) 40 Corpus bursae with spines forming radiating arc at the anterior edge (fig. 4d) .............. E. inouei Kawabe – Corpus bursae with spines not forming radiating arc ............................................................................... 41 41 Antrum with minute spines posteriorly .................................................................................................... 42 – Antrum without spines .............................................................................................................................. 43 42 Ductus bursae as long as corpus bursae; corpus bursae entirely spined ( Razowski, 1968: fig. 43 ) .............. .............................................................................................................................. E. charixantha (Meyrick) – Ductus bursae twice length of corpus bursae; corpus bursae lacking spines medially ( Razowski, 1968: fig. 46 ) ........................................................................................................................... E. eucalypta (Meyrick) 43 Corpus bursae entirely spined ................................................................................................................... 44 – Corpus bursae partially spined................................................................................................................... 45 44 Ductus bursae narrow, 1/4 width of corpus bursae; corpus bursae oblong (fig. 4e) .. E. kobeana Razowski – Ductus bursae broad, 1/3 width of corpus bursae; corpus bursae round ( Razowski, 2005: fig. 7 ) ............... ................................................................................................................................. E. yubariana Razowski 45 Corpus bursae with three spines ( Razowski, 1968: fig. 18 ) ....................................... E. crocina Razowski – Corpus bursae with more than ten spines ................................................................................................. 46 46 Ductus bursae with anterior portion as broad as posterior portion ........................................................... 47 – Ductus bursae with anterior portion broader than posterior portion ( Razowski, 1968: fig. 14 ) .................... ........................................................................................................................................... E. lata Razowski 47 Ductus bursae nearly as long as corpus bursae (fig. 4f) ................................................................................ ............................................................................................................ E. sanguisorbana (Herrich-Schäffer) – Ductus bursae longer than corpus bursae (fig. 4c) .................................................... E. citrinana Razowski