Taxonomic revision of the West ern Hemisphere checkered beetle genus Priocera KIRBY (Coleoptera, Cleridae, Clerinae) Author Opitz, Weston text Linzer biologische Beiträge 2021 52 2 1145 1255 journal article 10.5281/zenodo.5038854 0253-116X 5038854 Priocera castanea ( NEWMAN , 1838) ( Figs 85 , 88 , 114 ) Opilus castaneous NEWMAN, 1838: 380 . HOLOTYPE : ♀ . North America, Mount Pleasant, Ohio ( BMNH ). CORPORAAL 1950: 99 . KNULL 1951: 289 . EKIS (now OPITZ) 1975: 20. A d d i t i o n a l m a t e r i a l e x a m i n e d ( Fig. 88 ): In addition to the holotype , I examined 348 specimens from: The United States of America : Pennsylvania , Dauphin County, Rockville, 25-VII-1970 , H. B. Kirk; Philadelphia County , Manayunk , no other information ; Green County , Rutan , 6- VIII- 1966 , R. W. Surdick ; Fulton County , 0.2 km SE Barnes Gap , Sideling Hill Barrens , 19- VIII- 1996 , 245 m, light trap , W. Quin Metheny ; Clinton County , 5.4 km N Monument , 22-VII- 2011 , 347 m, P. Woods ; Allegheny County, no other information; New Jersey , Burlington County , Atsion , 10-VI-1945 , J. W. Green ; Middlesex County , Jamesburg ,?-IX-?, collector not noted ; Ocean County ,?- VII-1915 , collector not noted ; Burlington County , Medford Township , Lake Pine , 2-VII-1975 , D. C. Rentz ; idem, Tabernacle Turnpike , Oak Shade-Flyatt Road , 30-VI-1971 , blacklight trap , Howard P. Boyd ; Fulton County , 3 m N Cowans Gap Park , 10-VII-1988 , J. E. Wappes , Illinois , Will County , Joliet , 20-VII-1935 , collector not noted ; Putnam County , 8 km W Magnolia , 24-VII-1985 , J. Powell ; La Salle County , Starved Rock , 3-VII-1948 , Floyd G. Werner ; Fayette County , 2.7 km SW Vandalia , 22-VI-22-VII- 2016 , 168 m, C. Wetzel ; Saint Clair County , 1.1 km NNW East Saint Louis , 14-30-VI-2016 , S. Mils ; McLean County , 12-VII-1896 , A. B. Wolcott ; New York , Suffolk County , Yaphank , Long Island , 30-VII-1895 , A. C. Weeks ; Bronx County , Van Courtland Park , no other information ; Seneca County ,?- VIII- 1971 , Lenczy ; Tompkins County , Ithaca , Cornell Campus , 1- VIII- 1979 , R. & K. Gemmill ; Tompkins County , Groton , 20- VI-1956 , N. M. Downie ; Indiana , Porter County , Dunes State Park , 9-VII-1966 , M. N. Downie ; Tippecanoe County , W Lafayette , 24-VI-1977 , J. F. MacDonald ; Kansas , Johnson County , Overland Park Arboretum , nursery, 6-12-VI-2006 , canopy trap , A Cruz ; Wabaunsee County , Bolton Wildlife Area , near Paxico, NE access point, 22-V-2012 , Z. H. Falin ; Woodson County , Cross Timbers State Park , East Spillway Access , near lagoon, 8-11-VI-2010 , Z. H. Falin , K. T. Eldrege , S. R. Cave , J. Ortega-Blanco ; Douglas County , Lawrence , KU West Campus , 8-14-VII-1995 , Steve Lingafelter ; Pottawatomie County , Onaga , no other information available ; Labette County , Oswego , 28-VI-1962 , night lights, G. F. Hevel ; idem, 20-VI-1964 , under bark, G. H. Hevel ; Clark County , State Lake , 19- VIII- 2004 , Malaise trap , G. Salsbury ; Ohio , Scioto County , 3-VII-1949 , J. F. Smith ; Clinton County , 9-VII-1956 , collector not noted ; Clinton County , 8-VII-1956 , F. Moore ; Allen County , Lima , 16-VII-1990 , J. A. Green ; Auglaize County , 1 km NE Wapakoneta , 10-22-VII-2013 , C. Poe ; Hamilton County , Cincinnati , no other information ; Hocking County , 8- VIII- 1924 , J. N. Knull ; Michigan , Monroe County , 26-VI-1965 , Malaise trap , James Truchan ; Allegan County , Allegan State Forest , 3- VIII- 1972 , collector not noted ; Wayne County , Detroit , Schwarz ; Ingham County , East Lansing , 21-VII-1972 , Daniel K. Young ; Montana , Ravalli County , New Florence , 6-VII-1980 , Douglas Yanega ; Missouri , Randolph County , 10- VIII- 1975 , E. G. Riley ; Carter County , Pinewoods Lake , 22-VI-1987 , reared from dead Pinus echinata , T. C. MacRae ; Saint Louis County , 2.4 km E Maryland , heights, 27-VI-11-VII- 2016 , 193 m, L. Trevathan ; Cole County , 25- VII-1970 , Ramon Gass ; Boone County , 30-VI-1974 , P. Kinslowe ; idem, Columbia , 27-VII-1971 , W. S. Craig ; Jefferson County , Arnold , Bay Shore subdivision, 16-VII-1975 , S. E. Thewke ; Holt County , Mound City , 14-VII-1968 , Malaise trap , collector not noted ; Randolph County , Moberly , 11- VIII- 1972 , W. Bivin ; Washington D. C. , Glen Echo ,?- VII-1939 , J. C. Bridwell ; West Virginia , Greenbrier County , White Sulphur Springs ,?-VII-?, collector not noted ; Virginia , Roanoke County , Roanoke , 12-17-VI-1991 , E. Riley ; Rockbridge County , 2.1 km NE Buena Vista , 20-IV-17-VI- 2018 , D. Hetzel ; Maryland , Prince George’s County , Beltsville , 21- VIII- 1958 , H. P. Lanchester ; Montgomery County , Plummers Island , 13-VII-2002 , W. E. Steiner & J. M. Swearinger ; Talbot County , Wittman , 28-VII-1974 , W. E. Steiner ; Little , Bennett Regional park, Campground area , 25- VII- 2015 , 180 m, MV/UV light, S. M. Lingafelter ; Tennessee , Hamilton County , Signal Mountain , 22-VII-1921 , C. F. Dalman ; North Carolina , Craven County , 1 mile NE, Bachlor , 14-VI-1965 , H. A. Hespenheide ; Catawba County , Hog Hill 3-10-VI-1976 , R. Turnbow ; Currituck County , Currituck , 13-VI-1991 , R. W. Surdick ; Dare County , Roanoke Island , 16-VI-1983 , S. W. Gifford ; Wake County , 2 miles NW Raleigh , 19-VI-1964 , George A. Matuza ; Dare County , Manteo , 13-VII- 1983 , S. M. Gifford ; Moore County , Southern Pines , Durham County , Durham , no other information ; Moore County , Southern Pines , 7- VIII- 1912 , A. H. Manee ; South Carolina , Pickens County , Clemson , Clemson University , 24-V-1931 , Alfred Lutken ; Greenville County , Greenville , 26-VII-1974 , at lights, Richard S. Peigler ; Georgia , Thomas County , Thomasville , 12-V-1948 , F. Werner , W. Nutting ; Calhoun County , Cordrays Pond , 14-VI-1946 , P. W. Fattig ; Montgomery County , Uvalda area ,?- VI-1990 , R. Morris ; Henry County , Nail Road , 9-V-1995 , R. Beshear ; Jackson County , Hardman Forest , 10-15- VIII- 1975 , R. Turnbow ; Clinch County , 6.3 miles NE Edith on highway 177, emerged from 4-6 inch diameter dead trunk of Taxodium distichum during 21-30-IV-2008 , L. C. Rich collected trunk on 31-II-2006 ; Glynn County , Brunswick , 5-VI-1952 , D. G. Kissinger ; Fulton County , 3.6 km NNE Union City , 30-VI-27-VII-2016 , E. Dendy ; Douglas County , 1.8 km SE Austell , 8-22-VII-2016 , E. Dendy ; Chatham County , 0.9 km E Port Wentworth , 11-25-VI-2015 , S. Davis ; Green County , 3.9 km W Siloam , 3-17-VI-2015 , R. Willits ; Lamar County , Milner , 19-VI-1933 , T. L. Bissell ; idem, Milner ,?- VI-1990 , R. Morris ; Walton County , Monroe , 20-VI-1945 , P. W. Fattig ; Clarke County , 10-VI-1968 , R. T. Franklin ; DeKalb County , Dunwoody ,?-?-1955, E. F. Menhinick ; DeKalb County , Stone Mountain area , 2-VII-1971 , J. E. Wappes ; idem, Whitehall Forest , 25-30-VI-1976 , R, Turnbow ; Florida , Liberty County , Torreya State Park , 6-V-1989 , R. Turnbow ; Wakulla County , Apalachicola National Forest , 20-27-VI-2005 , Malaise in pine-oak savanna ; Duval County , Jacksonville , 20-V-1959 , P. H. Thompson ; Tampa , collection date not noted, Hubbard & Schwartz ; Putnam County , Crescent City , collection date not noted, Hubbard & Schwartz ; Saint Lucie County , Capron , collection date not noted, Hubbard & Schwartz ; Alachua County , Gainesville , 30-IV-1976 , light, Joe Schuh ; Highlands County , Archibold Biological Station , 19-IV-1976 , William Rosenberg ; idem, Sebring , 10-IV-1991 , R. Morris ; Mississippi , Marshall County , Wall Doxey State Park , 5-6-VI-1999 , R. L. Brown ; Noxubee County , Noxubee Wildlife Refuge , 16- VIII- 1984 , Soowon Cho ; George County , Mixon Lake , hillside, 9-10-IV-1999 , Clint Alien ; Smith County , 1 mile N Raleigh , 14-15-VI-1985 , R. Brown , G. Burrows ; Warren County , Vicksburg , 15-VI-2014 , Ricky Patterson ; Winston County , Tombigbee National Forest , 1-VI-1999 , J. A. MacGown ; Jackson County , Gulf Island National Sea., 3-6-VI- 1984 , B. Brown , G. Baker ; Kemper County , 2 miles N Scooba , 8- VIII- 1991 , T.L.Schiefer ; Webster County , 2 miles SE Cumberland , 12-VI-1990 , T. L. Schiefer ; Holmes County , Holmes County State Park , 8-VI-1994 , T. L. Schiefer ; Tishomingo County , Tishomingo State Park , 6-VI-1992 , blacklight, P. K. Lago ; Claiborne County , 3.6 miles W Port Gibson , 12-VII-1993 , T. L. Schiefer ; Harrison County , Gulfport , 25-VI-1938 , J. H. Roberts ; Oktibbeha County , Starkville , 15-VII-1971 , at artificial lights, W. H. Cross ; 6 miles SW Starkville , 11-VII-1984 , R. L. & B. B. Brown ; Rankin County , 4 miles NNE Brandon , 23-VI-1973 , W. H. Cross ; Forrest County , Hattiesburg ,?- V-1957 , B. & B. Valentine ; Alabama , Tuscaloosa County , Tuscaloosa , 9-VI-1952 , B. Valentine ; Lee County , Opelika , 23-VI-1979 , T. King ; Bibb County , Glades Preserve , 5-VI-2003 , R. L. Brown ; Lawrence County , Joe Wheeler State Park , 25-V-2004 , T. L.Schiefer ; Baldwin County , Bon Secour N. W. Refuge , 6-IX-1988 , L. Corpus & T. Schiefer ; Monroe County , Haynes Island Park, 26-29-V- 1995 , D. M. Pollock ; Dale County , Fort Rucker Mil. Res. Reservation , 8-17-V-2015 , R. Turnbow ; Walker County , Jasper , 9-XI-1978 , light trap , Tina King ; Clay County , Cheaha State Park , Cheaha Mountain , 10- VIII- 1985 , 2,400 feet , E. G. Riley ; Jackson County , 1.8 km WNW Hollytree , 13-VI- 2008 , UV trap , Androw , Karns , Grisham ; Shelby County , Helena , 8-VI-1957 , at light, H. R. Steeves ; Jefferson County , Birmingham , 12-VI-1956 , collector not noted ; Blount County , Highland Lake , 12-24-IV-2013 , light trap , T. King ; Mobile County , Mobile , collection date not noted, collector information not readable ; Grand Bay , 10-VI-1907 , H. P. Löding ; Lousiana , Natchitoches County , Kisatchie National Forest , 2 miles E Red Bluff , 17-18-VI-1988 , C. B. Barr ; West Feliciana Parish , 2 miles W Weyanoke , 26-VI-1987 , Tunica Hills , William D. Shepard ; Bossier Parish , Barksdale Air Force Base , 12-VI-1996 , D. M. Pollock ; Clairborne Parish , Lake Clairborne State Park , 6-VII-1983 , E. G. Riley ; Natchitoches Parish , Voweils Mill , 2-VI-1975 , M. Druckenbrod ; Arizona , Flagstaff , 15-VI-1904 , Pinus , J. L. Webb ; Oklahoma , Cleveland County , Norman , 16-VII-1975 , light trap , William D. Shepard ; Oklahoma County , Luthe r, 26-VI-1991 , hickory/oak forest, Chuck Harp ; Latimer County , 5 miles W Red Oak , 27-VI-1977 , K. Stephan ; Texas , Lamar County , Camp Maxey , 3.4 km S Powderly , 16-VI-20-VI-2009 , Malaise, J. C . Abbot ; Bastrop County , Smithville , Stengl Biological Station , 20-V-3-VI-2011 , T. H. Atkinson ; Hardin County .?- VII-1991 , F. B. Camors ; Nolan County , Lake Sweetwater area , 29-VI-1989 , P. Morris ; Brazos County , 5 km SW Wellborn , 23-VII-2016 , lights, V. Belov ; Sabine County , 8 miles E Hemphill , 24-V-7-VI-2008 , Lindgren / jug trap , E. Riley , B. Raber , & D. Heffern ; Van Zandt County , 0.3 km N Edom , 11-25-VII- 2016 , 157 m ; Nacogdoches County , Nacogdoches , 8-VI-1960 , Entomology class ; Walker County , Huntsville , 19-VI-1971 , T. Rush ; Colorado County , Columbus , collection date not noted, Hubbard & Schwartz ; Culberson County , Guadalupe National Park , Upper Dog Canyon , 14-VII-1975 , D. E. Foster ; Kent County , The Woodlands , 12-V-1978 , J. E. Wappes ; Hardin County , Southern Pine Beetle infested trees,?- VII-1971 , F. B. Camore , Jr .; Montgomery County , Woodlands , 23-V-1980 , J. E. Wappes ; idem, 1-8-VII-1978 , R. Turnbow. Specimens are deposited in: ACMT , AMNH , CSUC , FMNH , FSCA , KSUC , MEMU , RFMC , RHTC , SEMC , UMRM , USNM , UTBC , WFBM , and WOPC . D i a g n o s i s: Forebody, pterothorax, and legs castaneous; antenna testaceous; elytra tricolorous, mostly castaneous, with 3 yellow markings, one at elytral base near mesoscutellum, one in front of middle near epipleural margin, one behind middle near sutural margin, apical 1/3 rd testaceous. R e d e s c r i p t i o n: Size: Length 8.0 mm; width 2.0 mm. Form: As in Fig. 114 . Head: Frons wider than length of antennal pedicel; EW/FW 33/30. Thorax: Pronotal sides wrinkled, pronotum proper and pronotal arch finely punctate, pronotum proper with 2 tumescences ( PW /PL 85/130); elytral disc with basal tumescenses, middle of disc concave, disc sculptured with striate, asetiferous punctures that extend beyond basal 1/2, but do not reach elytral apex, latter narrowly rounded ( EL /EW 330/60); metathoracic femur gradually increasing in diameter to femoral apex. Abdomen: Male visible abdominal ventrite V deeply widely emarginate distally; aedeagus as in Fig. 85. V a r i a t i o n: Size: Length 5.0-9.0 mm; width 1.2-2.5 mm . The yellow elytral spot near the mesoscutellum may be faintly visible or absent. N a t u r a l h i s t o r y: Specimens were collected mostly during spring and summer months. One specimen emerged from a dead trunk of bald Cyprus [ Taxodium distichum (LINNAEUS) RICH. (Cupressaceae) ]. Another specimen was taken from the bark of a species of pine [ Pinus LINNAEUS (Pinaceae) ]. Specimens were also collected in a Malaise and light traps; one at 180 m , another at 731 m . Josef N. Knull reported ( KNULL 1951: 289 ) that this species was collected from white oak infested with Melittomma MURRAY (Lymexylidae) larvae and pitch pine infested with Dendroctonus frontalis ZIMMERMANN (Curculionidae) . He also indicates that, "adults are nocturnal and rarely found". N o t e s: EKIS (now OPITZ) (1975: 20) unnecessarily designated a lectotype for this species. It is clear from Newman’s description that he examined only one specimen .