Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales) Author Rio, Cédric Del Author Stull, Gregory W. Author Franceschi, Dario De text European Journal of Taxonomy 2020 2020-05-14 645 1 130 journal article 22093 10.5852/ejt.2020.645 1cd473d9-9d96-438f-bab4-29c5e56f82b8 3829651 Rhyticaryum fasciculatum Becc. Fig. 31.18–31.25 Material examined Specimen used for endocarp and fruit description INDONESIA • 12 Oct. 2004 ; Heu Her 4 ; K [ K000271837 ] . Description FRUIT. Epicarp sparsely strigose, with yellow simple hairs with granular ornamentation. Mesocarp thin. Length 13.0– 20.5 mm , width 10–15 mm , thickness 7–9 mm . ENDOCARP. Brown, length ca 20.5 mm , width ca 12.3 mm , thickness ca 9 mm . Keel surrounding the endocarp in the plane of symmetry. Apex truncate, slightly inflated and asymmetrical in lateral view; base rounded, symmetrical. Outer surface of the endocarp ridged, with ca four ridges longitudinally, the median ridge reaching the point of the base. Ridges rounded and thin, with a median channel, enclosing 15–21 polygonal areoles. Areols elongate longitudinally, with freely ending ridgelets, some ridgelets arbuscular. Primary vascular bundle positioned outside the endocarp wall. Endocarp wall 514–575 µm thick excluding ridges ( 751–989 µm thick including ridges). Endocarp wall with three cell layers: outermost layer with ca 14 rows of anticlinally oriented cells, cells 24.6–54.4 µm in length, followed by a layer with ca 8 rows of periclinally oriented cells, cells 18.1–25.1 µm in width; innermost layer with one row of periclinally oriented cells, cells 8.7–11.9 µm in width, lining the locule surface with regularly spaced and large papillae; papillae 7.2–21.0 µm (av. 13.9 µm ) in diameter, 710 papillae per 0.25 mm 2 .