Order Soricomorpha Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 220 311 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Crocidura Wagler 1832 Crocidura Wagler 1832 , Isis: 275 . Type Species: Sorex leucodon Herman 1780 Synonyms: Afrosorex Hutterer 1986 ; Heliosorex Heller 1910 ; Leucodon Fatio 1869 ; Paurodus Schulze 1897 ; Praesorex Thomas 1913 ; Rhinomus Murray 1860 . Species and subspecies: 172 species with 56 subspecies: Species Crocidura aleksandrisi Vesmanis 1977 Species Crocidura allex Osgood 1910 Species Crocidura andamanensis Miller 1902 Species Crocidura ansellorum Hutterer and Dippenaar 1987 Species Crocidura arabica Hutterer and Harrison 1988 Species Crocidura arispa Spitzenberger 1971 Species Crocidura armenica Gureev 1963 Species Crocidura attenuata Milne-Edwards 1872 Species Crocidura attila Dollman 1915 Species Crocidura baileyi Osgood 1936 Species Crocidura baluensis Thomas 1898 Species Crocidura batesi Dollman 1915 Species Crocidura beatus Miller 1910 Species Crocidura beccarii Dobson 1887 Species Crocidura bottegi Thomas 1898 Species Crocidura bottegoides Hutterer and Yalden 1990 Species Crocidura brunnea Jentink 1888 Subspecies Crocidura brunnea subsp. brunnea Jentink 1888 Subspecies Crocidura brunnea subsp. pudjonica Sody 1936 Species Crocidura buettikoferi Jentink 1888 Species Crocidura caliginea Hollister 1916 Species Crocidura canariensis Hutterer, Lopez-Jurado and Vogel 1987 Species Crocidura caspica Thomas 1907 Species Crocidura cinderella Thomas 1911 Species Crocidura congobelgica Hollister 1916 Species Crocidura crenata Brosset, Dubost, and Heim de Balsac 1965 Species Crocidura crossei Thomas 1895 Species Crocidura cyanea Duvernoy 1838 Species Crocidura denti Dollman 1915 Species Crocidura desperata Hutterer, Jenkins and Verheyen 1991 Species Crocidura dhofarensis Hutterer and Harrison 1988 Species Crocidura dolichura Peters 1876 Species Crocidura douceti Heim de Balsac 1958 Species Crocidura dsinezumi Temminck 1842 Species Crocidura eisentrauti Heim de Balsac 1957 Species Crocidura elgonius Osgood 1910 Species Crocidura elongata Miller and Hollister 1921 Species Crocidura erica Dollman 1915 Species Crocidura fischeri Pagenstecher 1885 Species Crocidura flavescens (I. Geoffroy 1827) Species Crocidura floweri Dollman 1915 Species Crocidura foetida Peters 1870 Subspecies Crocidura foetida subsp. foetida Peters 1870 Subspecies Crocidura foetida subsp. doriae Peters 1870 Subspecies Crocidura foetida subsp. kelabit Medway 1965 Species Crocidura foxi Dollman 1915 Species Crocidura fuliginosa Blyth 1855 Subspecies Crocidura fuliginosa subsp. fuliginosa Blyth 1855 Subspecies Crocidura fuliginosa subsp. dracula Thomas 1912 Species Crocidura fulvastra Sundevall 1842 Species Crocidura fumosa Thomas 1904 Species Crocidura fuscomurina Heuglin 1865 Species Crocidura glassi Heim de Balsac 1966 Species Crocidura gmelini ( Pallas 1811 ) Species Crocidura goliath Thomas 1906 Subspecies Crocidura goliath subsp. goliath Thomas 1906 Subspecies Crocidura goliath subsp. nimbasilvanus Hutterer 2003 Species Crocidura gracilipes Peters 1870 Species Crocidura grandiceps Hutterer 1983 Species Crocidura grandis Miller 1911 Species Crocidura grassei Brosset, Dubost, and Heim de Balsac 1965 Species Crocidura grayi Dobson 1890 Species Crocidura greenwoodi Heim de Balsac 1966 Species Crocidura harenna Hutterer and Yalden 1990 Species Crocidura hildegardeae Thomas 1904 Species Crocidura hilliana Jenkins and Smith 1995 Species Crocidura hirta Peters 1852 Species Crocidura hispida Thomas 1913 Species Crocidura horsfieldii Tomes 1856 Species Crocidura hutanis Ruedi and Vogel 1995 Species Crocidura ichnusae Festa 1912 Species Crocidura indochinensis Robinson and Kloss 1922 Species Crocidura jacksoni Thomas 1904 Species Crocidura jenkinsi Chakraborty 1978 Species Crocidura jouvenetae Heim de Balsac 1958 Species Crocidura katinka Bate 1937 Species Crocidura kivuana Heim de Balsac 1968 Species Crocidura lamottei Heim de Balsac 1968 Subspecies Crocidura lamottei subsp. lamottei Heim de Balsac 1968 Subspecies Crocidura lamottei subsp. elegans Hutterer 1986 Species Crocidura lanosa Heim de Balsac 1968 Species Crocidura lasiura Dobson 1890 Species Crocidura latona Hollister 1916 Species Crocidura lea Miller and Hollister 1921 Species Crocidura lepidura Lyon 1908 Species Crocidura leucodon (Hermann 1780) Species Crocidura levicula Miller and Hollister 1921 Species Crocidura littoralis Heller 1910 Species Crocidura longipes Hutterer and Happold 1983 Species Crocidura lucina Dippenaar 1980 Species Crocidura ludia Hollister 1916 Species Crocidura luna Dollman 1910 Species Crocidura lusitania Dollman 1915 Species Crocidura macarthuri St. Leger 1934 Species Crocidura macmillani Dollman 1915 Species Crocidura macowi Dollman 1915 Species Crocidura malayana Robinson and Kloss 1911 Species Crocidura manengubae Hutterer 1982 Species Crocidura maquassiensis Roberts 1946 Species Crocidura mariquensis A. Smith 1844 Subspecies Crocidura mariquensis subsp. mariquensis A. Smith 1844 Subspecies Crocidura mariquensis subsp. neavei Wroughton 1907 Subspecies Crocidura mariquensis subsp. shortridgei St. Leger 1932 Species Crocidura maurisca Thomas 1904 Species Crocidura maxi Sody 1936 Species Crocidura mindorus Miller 1910 Species Crocidura miya Phillips 1929 Species Crocidura monax Thomas 1910 Species Crocidura monticola Peters 1870 Species Crocidura montis Thomas 1906 Species Crocidura muricauda Miller 1900 Species Crocidura musseri Ruedi and Vogel 1995 Species Crocidura mutesae Heller 1910 Species Crocidura nana Dobson 1890 Species Crocidura nanilla Thomas 1909 Species Crocidura negligens Robinson and Kloss 1914 Species Crocidura negrina Rabor 1952 Species Crocidura nicobarica Miller 1902 Species Crocidura nigeriae Dollman 1915 Species Crocidura nigricans Bocage 1889 Species Crocidura nigripes Miller and Hollister 1921 Subspecies Crocidura nigripes subsp. nigripes Miller and Hollister 1921 Subspecies Crocidura nigripes subsp. lipara Miller and Hollister 1921 Species Crocidura nigrofusca Matschie 1895 Species Crocidura nimbae Heim de Balsac 1956 Species Crocidura niobe Thomas 1906 Species Crocidura obscurior Heim de Balsac 1958 Species Crocidura olivieri (Lesson 1827) Subspecies Crocidura olivieri subsp. olivieri Lesson 1827 Subspecies Crocidura olivieri subsp. anchietae Bocage 1889 Subspecies Crocidura olivieri subsp. bueae Heim de Balsac and Barloy 1966 Subspecies Crocidura olivieri subsp. cara Dollmann 1915 Subspecies Crocidura olivieri subsp. cinereoaenea Rüppell 1842 Subspecies Crocidura olivieri subsp. darfurea Thomas and Hinton 1923 Subspecies Crocidura olivieri subsp. giffardi de Winton 1898 Subspecies Crocidura olivieri subsp. guineensis Cabrera 1903 Subspecies Crocidura olivieri subsp. hansruppi Hutterer 1981 Subspecies Crocidura olivieri subsp. hedenborgiana Sundevall 1843 Subspecies Crocidura olivieri subsp. kivu Osgood 1910 Subspecies Crocidura olivieri subsp. manni Peters 1878 Subspecies Crocidura olivieri subsp. martiensseni Neumann 1900 Subspecies Crocidura olivieri subsp. nyansae Neumann 1900 Subspecies Crocidura olivieri subsp. occidentalis Pucheran 1855 Subspecies Crocidura olivieri subsp. odorata Leconte 1857 Subspecies Crocidura olivieri subsp. spurelli Thomas 1910 Subspecies Crocidura olivieri subsp. sururae Heller 1910 Subspecies Crocidura olivieri subsp. toritensis Setzer 1956 Subspecies Crocidura olivieri subsp. zuleika Dollman 1915 Species Crocidura orientalis Jentink in Weber 1890 Subspecies Crocidura orientalis subsp. orientalis Jentink in Weber 1890 Subspecies Crocidura orientalis subsp. lawuana Sody 1936 Species Crocidura orii Kuroda 1924 Species Crocidura palawanensis Taylor 1934 Species Crocidura paradoxura Dobson 1886 Species Crocidura parvipes Osgood 1910 Species Crocidura pasha Dollman 1915 Species Crocidura pergrisea Miller 1913 Species Crocidura phaeura Osgood 1936 Species Crocidura picea Sanderson 1940 Species Crocidura pitmani Barclay 1932 Species Crocidura planiceps Heller 1910 Species Crocidura poensis Fraser 1842 Species Crocidura polia Hollister 1916 Species Crocidura pullata Miller 1911 Species Crocidura raineyi Heller 1912 Species Crocidura ramona Ivanitskaya, Shenbrot, and Nevo 1996 Species Crocidura rapax G. Allen 1923 Subspecies Crocidura rapax subsp. rapax G. Allen 1923 Subspecies Crocidura rapax subsp. kurodai Jameson and Jones 1977 Subspecies Crocidura rapax subsp. lutaoensis Fang and Lee 2002 Subspecies Crocidura rapax subsp. tadae Tokuda and Kano 1936 Species Crocidura religiosa (I. Geoffroy 1827) Species Crocidura rhoditis Miller and Hollister 1921 Species Crocidura roosevelti Heller 1910 Species Crocidura russula (Hermann 1780) Subspecies Crocidura russula subsp. russula Hermann 1780 Subspecies Crocidura russula subsp. cintrae Miller 1907 Subspecies Crocidura russula subsp. osorio Molina and Hutterer 1989 Subspecies Crocidura russula subsp. peta Montagu and Pickford 1923 Subspecies Crocidura russula subsp. pulchra Cabrera 1907 Subspecies Crocidura russula subsp. yebalensis Cabrera 1913 Species Crocidura selina Dollman 1915 Species Crocidura serezkyensis Laptev 1929 Species Crocidura shantungensis Miller 1901 Subspecies Crocidura shantungensis subsp. shantungensis Miller 1901 Subspecies Crocidura shantungensis subsp. quelpartis Kuroda 1934 Species Crocidura sibirica Dukelsky 1930 Species Crocidura sicula Miller 1900 Subspecies Crocidura sicula subsp. sicula Miller 1900 Subspecies Crocidura sicula subsp. aegatensis Hutterer 1991 Subspecies Crocidura sicula subsp. calypso Hutterer 1991 Subspecies Crocidura sicula subsp. esuae Kotsakis 1984 Species Crocidura silacea Thomas 1895 Species Crocidura smithii Thomas 1895 Subspecies Crocidura smithii subsp. smithii Thomas 1895 Subspecies Crocidura smithii subsp. debalsaci Hutterer 1981 Species Crocidura somalica Thomas 1895 Species Crocidura stenocephala Heim de Balsac 1979 Species Crocidura suaveolens ( Pallas 1811 ) Species Crocidura susiana Redding and Lay 1978 Species Crocidura tanakae Kuroda 1938 Species Crocidura tansaniana Hutterer 1986 Species Crocidura tarella Dollman 1915 Species Crocidura tarfayensis Vesmanis and Vesmanis 1980 Species Crocidura telfordi Hutterer 1986 Species Crocidura tenuis Müller 1840 Species Crocidura thalia Dippenaar 1980 Species Crocidura theresae Heim de Balsac 1968 Species Crocidura thomensis Bocage 1887 Species Crocidura trichura Dobson in Thomas 1888 Species Crocidura turba Dollman 1910 Species Crocidura ultima Dollman 1915 Species Crocidura usambarae Dippenaar 1980 Species Crocidura viaria I. Geoffroy 1834 Species Crocidura virgata Sanderson 1940 Species Crocidura voi Osgood 1910 Species Crocidura vorax G. Allen 1923 Species Crocidura vosmaeri Jentink 1888 Species Crocidura watasei Kuroda 1924 Species Crocidura whitakeri de Winton 1897 Species Crocidura wimmeri Heim de Balsac and Aellen 1958 Species Crocidura wuchihensis Wang 1966 Species Crocidura xantippe Osgood 1910 Species Crocidura yankariensis Hutterer and Jenkins 1980 Species Crocidura zaphiri Dollman 1915 Species Crocidura zarudnyi Ognev 1928 Species Crocidura zimmeri Osgood 1936 Species Crocidura zimmermanni Wettstein 1953 Discussion: Eurasian species revised by Jenkins (1976) , Indomalayan and Philippine species by Heaney and Ruedi (1994) and Ruedi (1995), Chinese species by Jiang and Hoffmann (2001) . Phenetic and Phylogenetic relationships of African and Palearctic species studied by Butler et al. (1989) , Maddalena (1990) and McLellan (1994) , and of Asian and Indomalayan species by Heaney and Ruedi (1994) , Ruedi (1996), and Ruedi et al. (1998) . Karyotypes of SE Asian species described by Ruedi and Vogel (1995) , those of Mediterranean isls shrews by Vogel et al. (1990) , and of African shrews by Schlitter et al. (1999) . No formal subgenera are recognized here, although the type species of proposed subgenera are indicated. For synonyms see Allen (1939) , Heim de Balsac and Meester (1977) and Hutterer (1986a) .