A new fossil Eurypinae genus and sexual dimorphism in the representatives of Omineus (Coleoptera: Mycteridae) from Tertiary amber of Europe
Author
Alekseev, Vitalii Igorevich
0000-0003-4390-5443
Shirshov Institute of Oceanology, Russian Academy of Sciences, Nahimovskiy prospekt 36, 117997 Moscow, Russia. alekseew 0802 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4390 - 5443 & Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia.
alekseew0802@yahoo.com
Author
Bukejs, Andris
Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas 13, Daugavpils LV- 5401, Latvia.
Author
Pollock, Darren Andrew
Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA.
text
Zootaxa
2020
2020-07-29
4820
3
551
562
journal article
8966
10.11646/zootaxa.4820.3.8
f702f2bf-e817-4863-96f7-f20015f61de3
1175-5326
4398204
B8AAC629-F431-49E2-B15C-80CCC0CE070C
Europoeurypus inglaeso
sp. nov.
(
Figs. 1–5
)
Material examined
.
Holotype
: No. 6081 [
MAIG
] (
No. JD
8112 ex coll.
Jonas Damzen
), adult, apparently female.
Almost
complete beetle inclusion (tarsomeres present on right pro- and mesotarsus only, apical parts of all left tibiae missing) included in transparent, flat, oval, orange piece of amber with dimensions 24×14×
4 mm
.
Syninclusions
are represented by detrital particles and a few stellate trichomes of plant origin.
Type
strata
. Baltic amber from Eocene amber-bearing
Blaue Erde
deposits (mostly Bartonian age, as interpreted for extinct Central European resin-producing forests according to
Bukejs
et al
. 2019
).
Type
locality
.
Yantarny
settlement (formerly Palmnicken),
Sambian
(Samland)
Peninsula
,
Kaliningrad Region
,
Russia
.
Diagnosis
. As stated above, for the new genus.
Description
.
Body
relatively large-sized, elongate, slightly convex dorsally; dark, unicolorous. Dorsal surface evenly punctured and covered with fine, decumbent, conspicuous pubescence.
Measurements
: body length
10.2 mm
, maximum body width
3.3 mm
(elytra postmedially); head length about 1.0 mm, maximum width of head (across eyes)
1.8 mm
; pronotal length
1.7 mm
, maximum pronotal width
2.3 mm
; elytral length
7.5 mm
, maximum elytral width (in humeral area)
2.8 mm
.
Head
moderately narrowed posterior of eyes. Compound eyes protruding, transverse, without intrafacetal setae, widely separated, with entire and very slightly emarginate anterior margin; ommatidial facets distinct and fine (as large as punctures on head). Frons convex, minimum distance between eyes 2.8× as wide as longitudinal eye diameter measured laterally. Ocular (suborbital) grooves indistinct. Frontal furrows present, straight, subparallel. Labrum with anterior margin almost straight, bearing long setae. Mandibles bidentate apically. Terminal maxillary palpomere broadly triangular. Antennal insertion slightly hidden from above. Antennae 11-segmented, short, reaching posteriorly to near one-third of pronotum, pubescent. Antennomeres 2–10 cylindrical, about 2× as long as wide; scape and antennomere 11 wider than other antennomeres; scape subquadrate; antennomere 11 ovoid, pointed api-cally, about 1.5× as long as antennomere 10.
FIGURE 5.
Reconstruction of
Europoeurypus inglaeso
gen.
et
sp. nov.
from Baltic amber: habitus, dorsal view. Body length = 10.2 mm.
Thorax
. Pronotum transverse, 1.3× as wide as long, widest at middle. Pronotal base distinctly narrower than elytral base. Anterior and lateral margins of pronotum slightly rounded; posterior margin slightly sinuate. Lateral pronotal carina present, discernible in ventral view in two-thirds of pronotal length basally. Anterior and posterior angles rounded, not produced. Pronotal disc slightly convex, uneven, with pair of small symmetrical triangular depressions at base; pronotum densely and finely punctate, distance between punctures equal to 0.2–0.3× diameter of one puncture. Prosternum long anterior of procoxae. Procoxal cavities apparently open posteriorly. Elytra elongate, disc moderately convex; dorsal surface evenly and irregularly punctate and pubescent. Pubescence decumbent, long (equal to 5.0–6.0× distance between punctures); distance between punctures equal to 0.5–1.0× diameter of one puncture. Scutellar shield as long as wide, rounded, densely pubescent, pointed posteriorly. Mesocoxal cavities open. Elytral epipleura narrow, widest in basal part and then gradually narrowed posteriorly, extended to abdominal ventrite 2. Metaventrite convex, densely punctate.
Legs
slender. Procoxae subconical, contiguous. Meso- and metacoxae narrowly separated. Femora flattened. Tibiae straight, subequal in length to femora, with two short, equal in length spurs apically. Pro- and mesotarsi pen-tamerous. Penultimate tarsomere distinctly widened distally, flat, non-lobed. Tarsal claws swollen basally, without acute basal tooth, symmetrical. Ratio of mesotarsomere lengths 5: 3: 3: 4: 6.
Abdomen
with five visible abdominal ventrites. Ventrites 1–3 connate, ventrites 4–5 free; sutures between ven-trites 1–2 and 2–3 flat, sutures between ventrites 3–4 and 4–5 shelf-like. Anterior (intercoxal) process of ventrite 1 broadly triangular, pointed apically. All ventrites punctate, setose, without discernible sex patch(es). Apical margin of ventrite 5 rounded. Ratio of lengths of ventrites 1–5 3.5: 3.5: 3.5: 3.0: 2.5 (medially).
Etymology
. The specific epithet
inglaeso
is a Latin word combination of “in glaeso” (meaning “in amber”), used as a noun in apposition and created analogously to the widely used biological terms “
in vivo
” and “
in vitro
”.
Remarks
. The
holotype
apparently is female due to the lack of sex patch on abdominal ventrite(s).