Cave-dwelling Neobisiidae in South China Karst, with descriptions of twelve new species of Bisetocreagris (Pseudoscorpiones, Neobisiidae) from Guizhou Province Author Feng, Zegang 0000-0002-5809-5984 Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China https: // orcid. org / 0000 - 0002 - 5809 - 5984 & College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, P. R. China chenj @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0325 - 2532 & Key Laboratory of Zoological Systematics and Application, Hebei Basic Science Center for Biotic Interaction, College of Life sciences, Hebei University, Baoding, Hebei 071002, P. R. China dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031 * Corresponding author chenj@ioz.ac.cn Author Zhang, Feng 0000-0002-3347-1031 Key Laboratory of Zoological Systematics and Application, Hebei Basic Science Center for Biotic Interaction, College of Life sciences, Hebei University, Baoding, Hebei 071002, P. R. China dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031 * Corresponding author dudu06042001@163.com Author Chen, Jun 0000-0002-5809-5984 Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China https: // orcid. org / 0000 - 0002 - 5809 - 5984 & College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, P. R. China chenj @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0325 - 2532 chenj@ioz.ac.cn text Zootaxa 2023 2023-12-28 5395 1 1 77 https://www.mapress.com/zt/article/download/zootaxa.5395.1.1/52550 journal article 10.11646/zootaxa.5395.1.1 1175-5334 10438327 CCA38E8E-26E5-47D4-A27D-305554F6097F Key to the cave-dwelling species of Bisetocreagris of China 1 Pedicel of pedipalpal patella much shorter (about one third or one fourth of patella length)........................... 2 - Pedicel of pedipalpal patella much longer (about half or more than half of patella length)............................ 8 2 Chelal finger significantly shorter than hand with pedicel.................................. B. titania ( Mahnert, 2003 ) - Chelal finger significantly longer than hand with pedicel...................................................... 3 3 Carapace without eyes or eyespots........................................................................ 4 - Carapace with 2 eyes or eyespots......................................................................... 7 4 Rallum with 7 pinnate blades; femur length more than 4.90 ( ♀ ) mm, patella length more than 5.30 ( ♀ ) mm........................................................................................ B. gracilenta Gao et Zhang, 2017 - Rallum with 8–11 pinnate blades; femur length less than 4.00 ( ♂ , ♀ ) mm, patella length less than 4.00 ( ♂ , ♀ ) mm........ 5 5 Rallum with 11 pinnate blades; movable finger more than 3.30 ( ♂ , ♀ ) mm...................... B. longidigitata sp. nov. - Rallum with 8–9 pinnate blades; movable finger less than 2.30 ( ♂ , ♀ ) mm........................................ 6 6 Pedipalpal femur 7.5 (tritonymph) times longer than wide, patella 7.0 (tritonymph) times longer than wide, chela with pedicel 8.65 (tritonymph) times longer than wide.............................................. B. scaura ( Mahnert, 2003 ) - Pedipalpal femur 8.14‒8.81 ( ♂ , ♀ ) times longer than wide, patella 6.84–7.40 ( ♂ , ♀ ) times longer than wide, chela with pedicel 7.13‒8.02 ( ♂ , ♀ ) times longer than wide...................................................... B. brevis sp. nov. 7 Pedipalp without granulation; pedipalpal femur 8.6‒9.0 ( ♂ , ♀ ) times longer than wide, patella 9.1‒9.6 ( ♂ , ♀ ) times longer than wide..................................................................... B. juanxuae Mahnert and Li, 2016 - Pedipalp with granulation present on chelal hand; pedipalpal femur 7.14‒8.25 times longer than wide, patella 6.60‒7.72 times longer than wide........................................................................ B. grandis sp. nov. 8 Pedicel of pedipalpal patella obviously more than half the entire length of patella................... B. changbing sp. nov. - Pedicel of pedipalpal patella about half the entire length of patella............................................... 9 9 Carapace without eyes or eyespots....................................................................... 10 - Carapace with eyes or eyespots......................................................................... 13 10 Carapace with granulation present on lateral margin, pedipalp with obvious granular in all segments except chelal finger middle to the fingertip.......................................................................... B. aspera sp. nov. - Carapace without granulation, pedipalp patella without granulation............................................. 11 11 Pedipalpal femur more than 7.00 ( ♂ , ♀ ) times longer than wide, patella more than 6.00 ( ♂ , ♀ ) times longer than wide, chela with pedicel more than 7.2 ( ♂ , ♀ ) times longer than wide.......................... B. baozinensis Mahnert et Li, 2016 - Pedipalpal femur less than 5.30 ( ♂ , ♀ ) times longer than wide, patella less than 4.30 ( ♂ , ♀ ) times longer than wide, chela with pedicel 4.80 ( ♂ , ♀ ) times longer than wide................................................................ 12 12 Pedipalpal femur length 2.29 ( ♀ ) mm, patella length 2.28 ( ♀ ) mm, chela with pedicel length 3.75 ( ♀ ) mm................................................................................ B. maomaotou Gao, Wynne et Zhang, 2018 - Pedipalpal femur length 1.65‒1.68 ( ♂ , ♀ ) mm, patella length 1.42‒1.44 ( ♂ , ♀ ) mm, chela with pedicel length 2.49‒2.59 ( ♂ , ♀ ) mm............................................................................... B yanggong sp. nov. 13 Carapace with 2 eyes or eyespots........................................................................ 14 - Carapace with 4 eyes or eyespots........................................................................ 21 14 Pedipalpal femur less than 4.90 ( ♂ , ♀ ) times longer than wide, patella less than 3.60 ( ♂ , ♀ ) times longer than wide................................................................................. B. chuanensis Mahnert et Li, 2016 - Pedipalpal femur more than 4.90 ( ♂ , ♀ ) times longer than wide, patella more than 3.60 ( ♂ , ♀ ) times longer than wide.... 15 15 Pedipalpal femur more than 6.90 ( ♂ , ♀ ) times longer than wide, patella more than 6.00 ( ♂ , ♀ ) times longer than wide.... 16 - Pedipalpal femur less than 6.90 ( ♂ , ♀ ) times longer than wide, patella more than 6.00 ( ♂ , ♀ ) times longer than wide..... 17 16 Pedipalpal patella 6.07‒6.75 ( ♂ , ♀ ) times longer than wide, chela with pedicel 5.90‒6.71 ( ♂ , ♀ ) times longer than wide, chelal finger with 125–143 teeth................................................................. B. sanmen sp. nov. - Pedipalpal patella 7.84‒10.36 ( ♂ , ♀ ) times longer than wide, chela with pedicel 8.10–8.78 ( ♂ , ♀ ) times longer than wide, chelal finger with 163‒182 teeth.................................................... B. xiaoensis Liet Liu, 2017 17 The distance between trichobothrium sb and st longer than that of sb between b ................................... 18 - The distance between trichobothrium sb and st is somewhat equal to that of sb between b ........................... 19 18 Epistome rounded; pedipalp without modification, pedipalpal patella 4.69 ( ♂ ), 4.75 ( ♀ ) times longer than wide............................................................................... B. chinacavernicola ( Schawaller, 1995 ) - Epistome triangular; pedipalp with scale-like sculptured, pedipalpal patella 3.93‒4.40 ( ♂ ), 3.89‒4.41 ( ♀ ) times longer than wide................................................................. B. guangshanensis Gao et Zhang, 2017 19 The distance between trichobothrium st and t significantly shorter than 1/2 of sb between b , pedipalpal femur less than 5.90 ( ♂ , ♀ ) times longer than wide, patella less than 4.60 ( ♂ , ♀ ) times longer than wide....................... B. xishui sp. nov. - The distance between trichobothrium st and t significantly longer than 1/2 of sb between b , pedipalpal femur more than 6.10 ( ♂ , ♀ ) times longer than wide, patella more than 4.80 ( ♂ , ♀ ) times longer than wide............................... 20 20 Pedipalp with granulation present on chelal hand, trichobothrium it closer to fingertip than et , apex of coxae with 4–5 setae on each side............................................................................... B. chumi sp. nov. - Pedipalp without granulation, trichobothrium it -et at same level near fingertip, apex of coxae with 7–8 setae on each side.......................................................................................... B. jiangi sp. nov. 21 Pedipalp with granulation present on femur and chelal hand, pedipalpal femur less than 4.30 ( ♀ ) times longer than wide, patella less than 2.80 ( ♀ ) times longer than wide, chela with pedicel less than 3.50 ( ♀ ) times longer than wide................................................................................................. B. martii ( Mahnert, 2003 ) - Pedipalp with granulation present on chelal hand, pedipalpal femur more than 4.90 ( ♂ , ♀ ) times longer than wide, patella more than 3.40 ( ♂ , ♀ ) times longer than wide, chela with pedicel more than 4.00 ( ♂ , ♀ ) times longer than wide.............. 22 22 Pedipalpal femur more than 7.00 ( ♂ , ♀ ) times longer than wide, patella more than 6.00 ( ♂ , ♀ ) times longer than wide, chela with pedicel more than 6.50 ( ♂ , ♀ ) times longer than wide................................... B. tenuipalma sp. nov. - Pedipalpal femur less than 6.00 ( ♂ , ♀ ) times longer than wide, patella less than 5.00 ( ♂ , ♀ ) times longer than wide, chela with pedicel less than 5.50 ( ♂ , ♀ ) times longer than wide........................................................ 23 23 Pedipalpal femur more than 5.82–5.83 ( ♂ , ♀ ) times longer than wide, patella more than 4.34–4.67 ( ♂ , ♀ ) times longer than wide, chela without pedicel more than 4.91–5.05 ( ♂ , ♀ ) times longer than wide......... B. cavernarum Mahnert et Li, 2016 - Pedipalpal femur more than 5.08 ( ♀ ) times longer than wide, patella more than 3.63 ( ♀ ) times longer than wide, chela without pedicel more than 3.65 ( ♀ ) times longer than wide.............................................. B. ampla sp. nov.