A review of the worldwide distribution of Marenzelleria viridis, with new records for M. viridis, M. neglecta and Marenzelleria sp. (Annelida: Spionidae)
Author
Radashevsky, Vasily I.
A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, 17 Palchevsky Street, Vladivostok 690041, Russia.
Author
Pankova, Victoria V.
A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, 17 Palchevsky Street, Vladivostok 690041, Russia.
Author
Malyar, Vasily V.
A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, 17 Palchevsky Street, Vladivostok 690041, Russia.
Author
Cerca, José
Natural History Museum, University of Oslo, P. O. Box 1172, Blindern, NO- 0318 Oslo, Norway.
Author
Struck, Torsten H.
Natural History Museum, University of Oslo, P. O. Box 1172, Blindern, NO- 0318 Oslo, Norway.
text
Zootaxa
2021
2021-12-13
5081
3
353
372
journal article
2998
10.11646/zootaxa.5081.3.3
a8980f07-e2f6-440e-a3fb-5b26636382ee
1175-5326
5775688
0173F43D-38AB-4010-8B2A-1DFD5F118A2C
Marenzelleria neglecta
Sikorski & Bick, 2004
(
Figs 1
&
2
)
Marenzelleria neglecta
Sikorski & Bick, 2004: 264–268
, figs 2B, 3C, 5A–I, 6.
Syomin
et al.
2017: 977−979
, figs 2−5.
Blank & Bastrop 2009: 316-318
, fig. 1 (phylogeny).
Marenzelleria viridis
:
Maciolek 1984
(
Part
.)
: 51–55, fig. 2.
Bick & Burckhardt 1989: 239−241
, fig. 1, pls VII−VIII.
Bastrop
et al.
1995
: Baltic Sea population.
Bochert & Bick 1995: 764−772
, figs 2−12 (larval morphology).
Bochert 1996a: 58−68
, figs 1−31 (oogenesis); 1996b: 192−198, figs 1−27 (spermatogenesis).
Röhner
et al.
1996a
: Baltic Sea population.
Ysebaert
et al.
1996: 178−179
. Sikorski & Buzhinskaya 1998: 1118−1119, fig. 4.
Fide
Sikorski & Bick 2004
(
Part
.): 264. Not
Verrill 1873
.
?
Marenzelleria viridis
:
Cohen & Carlton 1995
: A4-1.
Thompson
et al
. 2000: 15
;
2013: 2290
.
Cohen
et al.
2005: 11
.
Ranasinghe
et al
. 2012: 646
.
Marenzelleria
cf.
viridis
:
Bick & Zettler 1997: 141−142
, figs 3, 4.
Fide
Sikorski & Bick 2004: 264
.
Marenzelleria
type
II:
Röhner
et al.
1996b: 281−283
.
Bastrop
et al.
1997: 125−127
;
1998: 98−102
.
Fide
Sikorski & Bick 2004: 264
.
Remarks.
Bastrop
et al.
’s (1997, 1998)
Marenzelleria
Type
II was found on the both sides of the North Atlantic and not formally named until
Sikorski & Bick (2004)
described it as a new species,
M. neglecta
. As noted above, Darss- Zingst-Boddenchain, Baltic Sea,
Germany
, was designated as the
type
locality of the species, although
M. neglecta
was recognized as alien in Europe, being introduced with ballast water from the Atlantic coast of
USA
(
Bastrop
et al.
1997
,
1998
).
Based on genetic data (allozyme electrophoresis and/or
16S
sequences),
Bastrop
et al.
(1997
,
1998
) documented the presence of
M. neglecta
(as
M.
Type
II) in the
United States
(Durham,
New Hampshire
; Chesapeake Bay,
Maryland
; Currituck Sound,
North Carolina
; Ogeechee River estuary,
Georgia
) and in
Canada
(Tuktoyaktuk Harbor,
Northwest Territories
) (
Fig. 2
). Notably, specimens of the only sample from
New Hampshire
(Durham, USNM 80485) were morphologically assigned to
M. viridis
by
Maciolek (1984
*),
Sikorski & Bick (2004)
, as well as one of us (VIR,
May 2000
).
Bastrop
et al
. (1997: 128)
sequenced the
16S
mitochondrial fragments of
three specimens
from this sample and noticed that “Contradictions between the results of morphological and genetic species determination still exist … DNA sequence analysis has shown beyond doubt that these animals [USNM 80485] belong to
Marenzelleria
Type
II [
M. neglecta
].”
Based on the morphology,
Sikorski & Bick (2004)
identified
M. neglecta
from San Francisco Bay (
California
,
USA
). After the description of
M. neglecta
, Leslie H. Harris
(
in litt
. 2 June,
5 July 2021
) re-identified as
M. neglecta
specimens initially reported as
M. viridis
from
California
,
USA
(see below Remarks for
M. viridis
). Moreover, Leslie H. Harris (
in litt
.
5 July 2021
) identified
M. neglecta
collected during the 2020 National Coastal Condition Assessment from Ilwaco marina, Columbia River mouth,
Washington
,
USA
(see VIR ID
24756 in
Table S4). Following these identifications, reports of
M. viridis
from San Francisco Bay by
Thompson
et al.
(2000
,
2013
) are also referred herein to
M. neglecta
. Complete information about records of
M. neglecta
from North America is given in Table S4.
Distribution in North America.
Atlantic coast: from
New Hampshire
south to
Georgia
,
USA
. Pacific coast:
Washington
and Northern
California
,
USA
. Arctic coast:
Northwest Territories
,
Canada
(
Fig. 2
).
*
Maciolek (1984: 52)
mistakenly exchanged the catalogue numbers for
two specimens
collected by
Jones, M. L.
in
Gloucester
(
Massachusetts
) on
16 May 1959
(
USNM 80483
), and
nine specimens
collected by
Long, C. D.
at
Fox Point
(
Durham
,
New Hampshire
) on
24 Jul 1962
(
USNM 80485
)
.