Systematics of Snakes Referred to Dipsas variegata in Panama and Western South America, with Revalidation of Two Species and Notes on Defensive Behaviors in the Dipsadini (Colubridae) Author CADLE, JOHN E. Author MYERS, CHARLES W. text American Museum Novitates 2003 2003-05-22 3409 1 48 http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29409%3C0001%3ASOSRTD%3E2.0.CO%3B2 journal article 10.1206/0003-0082(2003)409<0001:SOSRTD>2.0.CO;2 0003-0082 5373821 Dipsas nicholsi PANAMA : Colón : Agua Clara [about 9°11̍N, 79°41̍W; see footnote 10] ( Dunn, 1940 , 1949a ). Panama´: ‘‘Chagres Villages’’ [between Juan Mina and Madden Dam; see footnote 9] ( Dunn, 1940 , 1949a [1 head not examined for present study = ANSP 21907]). Madden Forest Preserve, 60–150 m [09°06̍N, 79°37̍W], KU 110310– 110314 (including cleared & stained, disarticulat­ ed, partially damaged skull of KU 110310). Madden Highway [probably in Madden Forest, see above], FMNH 217310. ‘‘Mid­basin of Chagres River and mouth of Pequeni River’’ [1933 type locality clarified herein as junction of Río Pequení with Río Boquerón (since 1936 = approximately mouth of Río Pequení near head of Madden Lake, about 20 km NNE Madden Dam), in upper drainage of the Río Chagres at 9°21̍N, 79°33̍W], MCZ 37884 ( holotype , head and neck only).