Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia Author Cárdenas-Bautista, Johann Stephens Author Parada-Alfonso, Jenny Andrea Author Carvajal‑Cogollo, Juan E. text Check List 2020 2020-07-03 16 4 821 839 http://dx.doi.org/10.15560/16.4.821 journal article 10.15560/16.4.821 1809-127X Onthophagus gr. clypeatus Blanchard, 1843 Figure 2B Materials examined. COLOMBIA • 1, 6.48 mm (mean); Meta , Villavicencio , Vereda Buenavista , “El Porvenir” Farm ; 04°08ʹ40.44″N , 073°41ʹ05.84″W ; 1207 m a.s.l. ; 25 Apr.2019 ; Cárdenas Johann and Parada Jenny leg.; wooded pasture; Bvpat4; UPTC-In-00084 . • 3, 6.48 mm (mean); Meta , Villavicencio , Vereda Buenavista , “El Porvenir” Farm ; 04°08ʹ42.33″N , 073°41ʹ09.20″W ; 1147 m a.s.l. ; 13 Oct. 2018 ; Cárdenas Johann and Parada Jenny leg.; riparian forest; Bvbrt7; UPTC-In-00085 . • 3, 6.48 mm (mean); Meta , Villavicencio , Vereda Buenavista , “El Porvenir” Farm ; 04°08ʹ35.25″N , 073°40ʹ50.28″W ; 1122 m a.s.l. ; 12 Oct. 2018 ; Cárdenas Johann and Parada Jenny leg.; high secondary forest; Bvbt5; UPTC-In-00086 . Identification. This species can be distinguished from other Onthophagus species by the prolonged clypeus on its anterior margin in the form of a “ T ”, slightly bidentate in the females; males with slightly angled horns, variable in size towards the base of the eyes; females with two conical protuberances towards the base of the eyes; and pronotum with a pair of small tubercles towards the midline that do not extend beyond the anterior border ( Zunino and Halffter 1997 ). Distribution. It is distributed throughout almost the entire country in Colombia , according to collection records. However, some differences between these species were observed. In the study area the species were found in all habitats between 650–1200 m a.s.l., and were very abundant (1954 individuals). However, a preference was observed for areas with forested vegetation (riparian forest relics and high secondary forest) where 1695 individuals were collected with some specimens reported for open areas such as wooded grasslands (259 individuals) and only seven individuals in the open grassland.