The genus Jeekelosoma Mauriès, 1985 - Moroccan cave millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) Author Enghoff, Henrik Author Reboleira, Ana Sofia P. S. text European Journal of Taxonomy 2019 2019-05-07 523 1 16 journal article 26904 10.5852/ejt.2019.523 7cefc0b8-0126-4022-a865-d2a8c0323c7c 2673211 0108D1A3-49B5-4EEE-8152-7714CDD17F3E Jeekelosoma heptarachne sp. nov. urn:lsid:zoobank.org:act: F43040BD-C8DA-474A-94CD-A9FB3B9202C6 Figs 5–8 Diagnosis A species of Jeekelosoma with 17 podous rings, with the ozopores borne on chimney-like outgrowths each provided with several macrosetae, with extremely long legs, and without a horizontal row of tubercles on podous rings 5–17. Etymology The name is a noun in apposition, composed of Greek ‘hepta’ (= seven) and ‘arachne’ (= spider). A male of the new species has 28 pairs of long, thin walking legs and thus (somewhat) resembles a row of seven spiders. Material examined Holotype MOROCCO • 3; Tazentout , cave Wit Tamdoun ; 9 Aug. 1971 ; C. Ribera leg.; MZB 73-2849 . Paratypes MOROCCO • 3 ♀♀ ; same collecting data data as for holotype; MZB 73-2849 , MZB 71-8589 , MZB 71-8590 • 1 ♂ ; in part mounted on two SEM stubs; MZB 71-8592 • 1 ♀ ; NHMD 302023 . Description Male SIZE. Body length ca 8 mm , max. width 0.63 mm , 19 ‘segments’ (17 podous rings + 1 apodous ring + telson). Fig. 5. Jeekelosoma heptarachne sp. nov. , paratype, female (NHMD 302023). Scale bar = 1 mm. HEAD. As in J. abadi , but densely covered by long setae. Length of antennae 3.1 × max. body width. Relative length of antennomeres 2–8: 18 / 20 / 17 / 16 / 20 / 7 / 2.3. COLLUM. As in J. abadi , but setae much longer. MIDBODY RINGS ( Fig. 6 A–C). Very narrow/elongated, length of metazona equaling maximum width. Body extremely narrow in anterior part, maximum width attained a little behind midbody; from there until and including ring 18 (the apodous ring in front of telson) body parallel-sided. Surface of metazona with very pronounced microcellulate structure, borders between cuticular scutes raised. Metazona with diameter increasing towards rear margin and with a clear constriction at ca ¹/3 of its length, measured from rear margin; posterior margin of rings finely crenulate but without a proper limbus. Each metazona with three rows of long, rather stout setae borne on distinct tubercles; each row on midbody rings with ca 10 setae; anterior row curving posteriad laterally; intermediate row situated just behind constriction. Metazonal setae much longer than in J. abadi (cf. Fig. 2 ). No horizontal row of tubercles, but lateral parts of anterior row appearing as oblique row of tubercles in lateral view. OZOPORES ( Fig. 6 G–H). On cylindrical peritremata, with 4–5 long, stout setae in a group anterodorsally to pore. LEGS ( Fig. 6C , E–F). Length 2.25 × max. body width. Relative length of podomeres: prefemur: 12 / femur: 30 / postfemur: 7 / tibia: 11 / tarsus: 35 / claw: 2.4. Femora clearly longer than body diameter, no distal swelling. No scopulae. TELSON. As in J. abadi . GONOPODAL APERTURE ( Fig. 6D ). Transverse elliptical, with a small median process originating on posterior margin. GONOPODS ( Figs 7–8 ). Coxa ( cx ) slightly convex on anterior surface; prefemoral part ( prf ) slightly more than half as long as acropodite; mesal acropodital process ( map ) a simple broad, dorso-ventrally flattened structure; solenophore ( sph ) as long as map , slender, apically with a subrectangular lamella ( sph‑a ) with distal corners drawn out into long processes, but without a terminal appendage; solenomere ( slm ) slender, simple, largely concealed within sph . Female Non-sexual characters as in male, except: SIZE. Body length 10–13 mm , max. width 0.58–0.80 mm . ANTENNAE. Length in largest female 2.0 × max. body width. Relative length of antennomeres 2–8: 18 / 18 / 16 / 16 / 21 / 7 / 3. LEGS. Length in largest female 1.8 × max. body width. Relative length of podomeres: prefemur: 13 / femur: 32 / postfemur: 8 / tibia: 11 / tarsus: 34 / claw: 2.4. Distribution and habitat Known only from “Tazentout, cave Wit Tamdoun”. This cave ( Bichain 2008 ) is also home to a rove beetle Domene cantonsi Español, 1972 known only from this and two further Moroccan caves ( Hernando 2007 ), and to an endemic North African terrestrial isopod, Trichoniscus soloisensis Vandel, 1959 ( Cruz 1991 ) .