Redescription of Heterocerus fausti Reitter, 1879, bona species (Coleoptera, Heteroceridae)
Author
Sazhnev, Alexey S.
text
Zootaxa
2018
2018-06-28
4441
3
597
600
journal article
29788
10.11646/zootaxa.4441.3.12
58c8ec85-77bc-49f4-ada7-3c923d1ec269
1175-5326
1301560
B2D326BE-54E6-4E6E-9DF6-00220F04EC41
Heterocerus fausti
Reitter, 1879
(
Figs. 1, 2B, 3
)
Heterocerus fausti
Reitter, 1879
: 546
stat. nov.
Type
locality.
Azerbaijan
:
Baku
.
Type
material examined.
Syntypes
: “
Heterocerus Fausti
Rtt
” [Reitter's handwriting], “Caucasus” [red label], “к.
А. Яковлева” [cоllеctiоn оf А. Jаcоvlеv—in Russiаn], 2♂♂ (ZISP); “Bаku Fаust.”, gоldеn squаrе plаtе, “
Heterocerus Fausti
Reitt.
” [Reitter's handwriting], 2♀♀ [one specimen without head] (ZISP).
Additional material examined.
9 exs.
TURKMENISTAN
.
Balkan
Region
(
Velayat
):
Uzun-Ada
(now the territory of
Hazar Nature Reserve
),
15.04.1895
, leg.
Varentsov
, 7 exs. (
ZISP
)
;
Turkmenbashi (
Krasnovodsk
), 0 6.1900, leg.
P. Maksimovitsh
, 1 ex. (
ZISP
)
; Esenguly, 3.07.1932, unknown collector, 1 ex. (IBIW).
Diagnosis.
Body size 3.8–5.0 mm. Body moderately elongated, dorsal-ventrally compressed (
Fig. 1
). Males and females do not differ. Aedeagus as in
Fig. 3
.
FIGURE 1–4.
Heterocerus fausti
Reitter, 1879
, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers
Heterocerus
spp. 2A—
H. parallelus
Gebler, 1830
, South Kazakhstan; 2B—
H. fausti
, Turkmenistan; 2C—
H. fossor
Kiesenwetter, 1843
, Russia, Dagestan; 2D—
H. flexuosus
Stephens, 1828
, Kazakhstan; 2E—
H. persicus
Mascagni, 1989
, Iran. Male genitalia of
Heterocerus fausti
, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens
Heterocerus
spp. 4A—
H. parallelus
, South Kazakhstan; 4B—
H. fausti
, Caucasus
; 4C—
H. fossor
, Russia, Dagestan; 4D—
H. flexuosus
, European part of Russia (Litovkin
et al
., 2013), 4E—
H. persicus
, Iran. Scale bar—0.5 mm.
FIGURE 5.
Distribution of
Heterocerus fausti
Reitter, 1879
. Red circles—studied material.
Heterocerus fausti
is similar in body form and drawing of elytra on
Heterocerus parallelus
but can be easily distinguished by the shape of its aedeagus (
Fig. 4A–B
) and smaller size.
Redescription.
Color.
Ground color is yellow or red-yellow; a combination of black-brown blotches on the pronotum, and brown blotches of elytra and abdomen form a characteristic pattern. Head and labrum are black-brown. Antennae completely yellow. Pronotum is black-brown with a more or less wide yellow edging on the sides and narrow yellow stripe along the front edge. Scutellum is black-brown. Drawing of elytra characteristic of
flexuosus
species-group sensu
Charpentier (1965)
. Yellow pattern of each elytron normally forms edging, S-shaped basal mark, M-shaped medial mark, apical spot and suture stripe. Dark pattern of the elytra brownish almost blending with the yellow background. Prosternum black-brown with yellow episternae and yellowish front margin. Mesepisternum and metepisternum blackbrown. Abdomen black-brown with yellow pattern in form of edging. Legs, except dark metacoxa, yellow.
Head.
Clypeus transversal, widely incised on the front edge. Antennae 11-segmented with 7-segment club. Hairs whitish. Puncturing uniform.
Thorax.
Pronotum not bordered basally; slightly narrower or comparable in width with base of elytra in females, and slightly wider than elytra base in males. Hairs on the disc are double: comprise dense, short, pressed, and long and standing hairs; whitish. Puncturing very dense, fine and uniform.
Elytra covered with erect hairs; hair length is comparable with the length of few first spines of mesotibia; hairs slightly longer along sides and near the elytral apex; hairs whitish.
Legs of structure typical for family, covered with whitish hairs. Claws of tarsus noticeably longer or equal to the last tarsomere (
Fig. 2
).
Abdomen
. Abdomen covered with whitish hairs. Postmetacoxal lines absent.
Remarks.
The records of
Heterocerus fausti
for
Uzbekistan
(
Jacobson, 1913
;
Zaitsev, 1946
) are not confirmed. Specimen from
Uzbekistan
(Qamashi) signed of F.A. Zaitsev as “
Heterocerus fausti
” is
Heterocerus heydeni
Kuwert, 1890
.
Ecology.
The long claws are probably an adaptation to the psammophilous way of life.
Distribution.
Pre-Caspian Region including Caucasus:
Azerbaijan
, West
Turkmenistan
(
Fig. 5
).