<document id="2A412D3E09BB7C4030B7D0A2637891EE" ID-CLB-Dataset="27521" ID-DOI="10.11646/zootaxa.4576.1.4" ID-GBIF-Dataset="a169fa80-80f0-4792-8a01-69b3bac059db" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624686" ID-ZooBank="A521DFD3-564F-4A15-8F59-9E946F169C4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272896246" checkinUser="plazi" docAuthor="Li, Bingxue, Zhang, Yuan &amp; Chen, Hongwei" docDate="2019" docId="03D787DCFFA5FFDAC69C5C8BFA22FB0D" docLanguage="en" docName="zootaxa.4576.1.4.pdf" docOrigin="Zootaxa 4576 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stegana coleoptrata Scopoli 1763" docType="treatment" docVersion="8" lastPageNumber="88" masterDocId="FFEEFFA4FFA2FFDDC60B5F7FFFBEFFD3" masterDocTitle="Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae)" masterLastPageNumber="108" masterPageNumber="81" pageNumber="88" updateTime="1698716517574" updateUser="ExternalLinkService">
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<mods:title id="D7C6D849AAFE0E7EAD53EF44C26DF51C">Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae)</mods:title>
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<mods:namePart id="3BD18145554CC8CECF7C3866DEF7C18F">Li, Bingxue</mods:namePart>
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<mods:namePart id="26A72A1CB193BFDBE0170BF4F47B159B">Zhang, Yuan</mods:namePart>
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<heading id="D08981A6FFA5FFDAC69C5C8BFD8BFBDD" bold="true" box="[151,565,1012,1038]" fontSize="11" level="1" pageId="7" pageNumber="88" reason="1">
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<taxonomicName id="4C7E4D49FFA5FFDAC69C5C8BFE3BFBDD" ID-CoL="8F4SH" authorityName="Scopoli" authorityYear="1763" box="[151,389,1012,1038]" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="coleoptrata">
<emphasis id="B90AEAD8FFA5FFDAC69C5C8BFE3BFBDD" bold="true" box="[151,389,1012,1038]" italics="true" pageId="7" pageNumber="88">Stegana coleoptrata</emphasis>
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species group
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<taxonomicName id="4C7E4D49FFA5FFDAC69C5B44FEDDFB83" ID-CoL="8F4SH" authorityName="Scopoli" authorityYear="1763" box="[151,355,1082,1104]" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="coleoptrata">
<emphasis id="B90AEAD8FFA5FFDAC69C5B44FEDDFB83" box="[151,355,1082,1104]" italics="true" pageId="7" pageNumber="88">Stegana coleoptrata</emphasis>
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species group: 
<treatmentCitation id="0ADF10DBFFA5FFDAC40A5B45FCB5FB83" author="Lastovka, P. &amp; Maca, J." box="[513,779,1082,1104]" page="8" pageId="7" pageNumber="88" year="1982">
<bibRefCitation id="EFEF4B3BFFA5FFDAC40A5B45FD4CFB83" author="Lastovka, P. &amp; Maca, J." box="[513,754,1082,1104]" pageId="7" pageNumber="88" pagination="1 - 38" refId="ref10439" refString="Lastovka, P. &amp; Maca, J. (1982) European and North American species of the genus Stegana (Diptera, Drosophilidae). Annotationes Zoologicae et Botanicae, 149, 1 - 38." type="journal article" year="1982">Laštovka &amp; Máca 1982</bibRefCitation>
: 8
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.
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<paragraph id="8BC136CAFFA5FFDAC69C5B02FAE6FB45" blockId="7.[151,1436,1149,1246]" box="[151,1368,1149,1174]" pageId="7" pageNumber="88">
<emphasis id="B90AEAD8FFA5FFDAC69C5B02FEAFFB45" bold="true" box="[151,273,1149,1174]" pageId="7" pageNumber="88">Diagnosis.</emphasis>
Surstylus submedially strongly curved inward, apically with 1 small prensiseta (
<figureCitation id="13452A4FFFA5FFDAC2925B02FAB1FB45" box="[1177,1295,1149,1174]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStart-6="FIGURE 9" captionStartId-0="11.[151,250,1191,1213]" captionStartId-1="12.[151,250,1890,1912]" captionStartId-2="13.[151,250,1301,1323]" captionStartId-3="14.[151,250,1577,1599]" captionStartId-4="15.[151,250,1814,1836]" captionStartId-5="16.[151,250,1193,1215]" captionStartId-6="17.[151,250,1567,1589]" captionTargetBox-0="[177,1402,220,1147]" captionTargetBox-1="[171,1426,379,1863]" captionTargetBox-2="[175,1403,235,1257]" captionTargetBox-3="[181,1397,447,1542]" captionTargetBox-4="[174,1407,723,1768]" captionTargetBox-5="[178,1407,208,1145]" captionTargetBox-6="[180,1398,566,1536]" captionTargetId-0="figure@11.[151,1436,193,1170]" captionTargetId-1="figure@12.[151,1436,339,1868]" captionTargetId-2="figure@13.[150,1436,193,1279]" captionTargetId-3="figure@14.[151,1435,411,1573]" captionTargetId-4="figure@15.[151,1436,697,1793]" captionTargetId-5="figure@16.[150,1436,193,1172]" captionTargetId-6="figure@17.[151,1436,538,1565]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="17" captionText-0="FIGURE 3. Stegana (Steganina) longifibula Takada, 1968, male terminalia: A. Epandrium (epand), surstylus (sur) and cercus (cerc) (lateral view); B, C. Hypandrium (hypd), parameres (pm), aedeagus (aed) and aedeagal apodeme (aed a) (ventral and lateral views); D, E. Gonopods (ventral and lateral views); F. Tenth sternite (ventral view). Scale lines = 0.1 mm." captionText-1="FIGURE 4. Stegana (Steganina) toyaensis Okada &amp; Sidorenko, 1992, male terminalia: A. Epandrium, surstylus and cercus (lateral view); B. Surstylus (inner view); C, D. Hypandrium, parameres, aedeagus and aedeagal apodeme (ventral and lateral views); E, F. Gonopods (ventral and lateral views); G. Tenth sternite (ventral view); H. Epiproct and cercus (dorsal view); I. Hypoproct (ventral view); J. Eighth sternite (ventral view). Scale lines = 0.1 mm." captionText-2="FIGURE 5. Stegana (Steganina) latitabula sp. nov., male terminalia: A. Epandrium, surstylus and cercus (lateral view); B. Surstylus (inner view); C, D. Hypandrium, parameres, aedeagus and aedeagal apodeme (ventral and lateral views); E, F. Gonopods (ventral and lateral views); G. Tenth sternite (ventral view). Scale lines = 0.1 mm." captionText-3="FIGURE 6. Stegana (Steganina) panda sp. nov., male terminalia: A. Epandrium, surstylus and cercus (lateral view); B. Surstylus (inner view); C, D. Hypandrium, parameres, aedeagus and aedeagal apodeme (ventral and lateral views); E, F. Gonopods (ventral and lateral views); G. Tenth sternite (ventral view). Scale lines = 0.1 mm." captionText-4="FIGURE 7. Stegana (Steganina) pinguifoliacea sp. nov., male terminalia: A. Epandrium, surstylus and cercus (lateral view); B, C. Hypandrium, parameres, aedeagus and aedeagal apodeme (ventral and lateral views); D, E. Gonopods (ventral and lateral views); F. Tenth sternite (ventral view). Scale lines = 0.1 mm." captionText-5="FIGURE 8. Stegana (Steganina) spatulata sp. nov., male terminalia: A. Epandrium, surstylus and cercus (lateral view); B. Surstylus (inner view); C, D. Hypandrium, parameres, aedeagus and aedeagal apodeme (ventral and lateral views); E, F. Gonopods (ventral and lateral views); G. Tenth sternite (ventral view). Scale lines = 0.1 mm." captionText-6="FIGURE 9. Stegana (Steganina) stachydifolia sp. nov., male terminalia: A. Epandrium, surstylus and cercus (lateral view); B. Surstylus (inner view); C, D. Hypandrium, parameres, aedeagus and aedeagal apodeme (ventral and lateral views); E, F. Gonopods (ventral and lateral views); G. Tenth sternite (ventral view). Scale lines = 0.1 mm." httpUri-0="https://zenodo.org/record/2624692/files/figure.png" httpUri-1="https://zenodo.org/record/2624694/files/figure.png" httpUri-2="https://zenodo.org/record/2624696/files/figure.png" httpUri-3="https://zenodo.org/record/2624698/files/figure.png" httpUri-4="https://zenodo.org/record/2624700/files/figure.png" httpUri-5="https://zenodo.org/record/2624702/files/figure.png" httpUri-6="https://zenodo.org/record/2624704/files/figure.png" pageId="7" pageNumber="88">Figs 3–9A</figureCitation>
or B).
</paragraph>
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Only those characters that depart from the description of the 
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<emphasis id="B90AEAD8FFA5FFDAC5745BDEFBBEFB6A" box="[895,1024,1185,1209]" italics="true" pageId="7" pageNumber="88">coleoptrata</emphasis>
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species group are provided here for the sake of brevity. For more detailed description see 
<bibRefCitation id="EFEF4B3BFFA5FFDAC4E55BBAFC46FB0D" author="Lastovka, P. &amp; Maca, J." box="[750,1016,1221,1246]" pageId="7" pageNumber="88" pagination="1 - 38" refId="ref10439" refString="Lastovka, P. &amp; Maca, J. (1982) European and North American species of the genus Stegana (Diptera, Drosophilidae). Annotationes Zoologicae et Botanicae, 149, 1 - 38." type="journal article" year="1982">Laštovka &amp; Máca 1982</bibRefCitation>
; 
<bibRefCitation id="EFEF4B3BFFA5FFDAC20F5BB9FB7DFB0D" author="Hu, Y. G. &amp; Toda, M. J." box="[1028,1219,1221,1246]" pageId="7" pageNumber="88" pagination="151 - 160" refId="ref10331" refString="Hu, Y. G. &amp; Toda, M. J. (1994) The Stegana (Steganina) coleoptrata species-group (Diptera, Drosophilidae), with descriptions of two new species and new records from eastern Palearctic region. Japanese Journal of Entomology, 62, 151 - 160." type="journal article" year="1994">Hu &amp; Toda 1994</bibRefCitation>
; 
<bibRefCitation id="EFEF4B3BFFA5FFDAC2C55BBAFA27FB0D" author="Bachli, G. &amp; Vilela, C. R. &amp; Escher, S. A. &amp; Saura, A." box="[1230,1433,1221,1246]" pageId="7" pageNumber="88" pagination="362" refId="ref9789" refString="Bachli, G., Vilela, C. R., Escher, S. A. &amp; Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. In: Fauna Entomologica Scandinavica. Fol. 39. Brill, Leiden, New York, pp. 362." type="book chapter" year="2004">
Bächli 
<emphasis id="B90AEAD8FFA5FFDAC3155BB9FAF1FB0E" box="[1310,1359,1221,1245]" italics="true" pageId="7" pageNumber="88">et al</emphasis>
. 2004
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.
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