A New Species Of The Genus Ramuliseta (Diptera, Ctenostylidae) From Madagascar, With A Key To Species
Author
Korneyev, V. A.
text
Vestnik Zoologii
2015
2015-12-01
49
6
489
496
http://dx.doi.org/10.1515/vzoo-2015-0060
journal article
10.1515/vzoo-2015-0060
2073-2333
6452665
Ramuliseta
Keiser, 1951
Keiser, 1951: 119
;
Hennig, 1960: 323
;
Steyskal, 1980: 556
;
McAlpine, 1990: 365
, 369;
Barraclough, 1994: 6
,
1995: 135
,
1998: 115
;
Korneyev, 2001: 48
;
2006: 63
,
2010: 963
, 965;
Rafael et al., 2009: 63
, 64;
Câmara & Rafael, 2013: 147
;
Pereira-Colavite & Mello, 2014: 219
.
Type
species:
Ramuliseta palpifera
Keiser, 1951
(by original designation).
Diagnosis. The flies of the genus
Ramuliseta
, similarly to the other Higher
Tephritoidea
(sensu
Korneyev, 1999
) have pictured wings and telesopic ovipositor of the females. Similarly to the other ctenostylids, they can be easily distinguished by the combination of lacking ocelli, vestigial proboscis, incomplete vein Sc, cell cup closed by arcuate crossvein without triangular lobe along vein CuA
2
+A
1
, strongly dimorphic arista (dendritical in females, simple and short pubescent in males), transverse prosternum, proepisternum bare or at most with one fine seta, spiracles round and open, without marginal fringe, anterior notopleural seta lacking, male genitalia with surstyli indistinguishable, without prensisetae, phallus spimple, tubular, with reduced glans, and female with aculeus simple, tubular, non-sclerotized, with small, blunt and short setulose, non-piercing cercal unit. Flies of the genus
Ramuliseta
have well developed palp as in species of Neotropical
Furciseta
Aczél, 1956
,
Oriental
Nepaliseta
Barraclough, 1995
and
Sinolochmostylia
Yang, 1995
differing from them all by veins R
2+3
and R
4+5
subparallel: distance along costal margin between R
2+3
and R
4+5
less than twice distance between R
4+5
and M (strongly divergent, with R
2+3
conspicuously bowed anterobasally in
Furciseta
,
Nepaliseta
, and
Sinolochmostylia
). The species of
Ramuliseta
differ from
Furciseta
also by having frons of males unmodified, not strongly narrowed (sexually dimorphic in
Furciseta
very narrow in males and unmodified in females) and vein M developed in distal half of cell bm (vein M lacking basally of crossvein bm-cu in
Furciseta
). The species of
Ramuliseta
differ from
Nepaliseta
and
Sinolochmostylia
by having flagellomere 1 of females elongate, with narrowed apical projection, and arista with multiple branching (in
Nepaliseta
and
Sinolochmostylia
, flagellomere 1 of females almost spherical, without apical projection, and arista extremely modified, disk-like).
The genus was described by
Barraclough (1994)
in details, and I do not repeat the description here.
D i s t r i b u t i o n. Afrotropical (
Nigeria
and
Tanzania
to
South Africa
;
Madagascar
) and Oriental Region (Lesser Sunda Islands).
The genus now includes 3 species: Palaeotropical
Ramuliseta palpifera
Keiser, 1951
(=
R. lindneri
Keiser, 1952
), Afrotropical
R. madagascariensis
Hennig, 1960
and
R. dolini
sp. n.
(
Madagascar
). The Oriental
R. thaica
Korneyev, 2001
belongs elsewhere, either to
Sinolochmostylia
or
Nepaliseta
(Korneyev, in prep.).
Key to species of
Ramuliseta
1. Smaller (wing shorter than
5.5 mm
) dark brown flies; wing with brown wing pattern; cell R
4+5
entirely brown in apical half (
fig. 1
,
3
). Mainland sub-Saharan Africa,
Lesser Sunda Islands
. ................................. .....................................................................................................................
Ramuliseta palpifera
(
Keiser, 1951
)
— Larger (wing shorter than
6.5 mm
) yellow flies; wing with yellow wing pattern; cell R
4+5
with one or two hyaline areas in apical half (
fig. 1,
1
,
2
).
Madagascar
..................................................................................... 2.
2. Wing apex hyaline; cell r
4+5
distal of crossvein dm-cu with two hyaline areas separated by rounded brown spot between them (
fig. 1
,
2
). ......................................
Ramuliseta madagascariensis
Hennig, 1961
— Wing apex yellow; cell r
4+5
distal of crossvein dm-cu with one large hyaline area and apical yellow band reaching its apex (
fig. 1,
1
). ..........................................................................................
Ramuliseta dolini
sp. n.