Vladimiretskia nathani gen. et sp. nov. (Diptera, Cecidomyiidae: Porricondylinae, Asynaptini, Vladimiretskiana subtr. nov.) from Eocene Rovno amber with description of their eggs Author Fedotova, Zoya A. 0000-0002-8888-5979 All-Russian Institute of Plant Protection of the Russian Academy of Sciences, 196608 Saint Petersburg - Pushkin, Russia; https: // orcid. org / 0000 - 0002 - 8888 - 5979 zoya-fedotova@mail.ru Author Vasilenko, Dmitry V. 0000-0002-4827-7290 Laboratory of Arthropods, A. A. Borissiak Paleontological Institute of the Russian Academy of Sciences, 117647 Moscow, Russia; https: // orcid. org / 0000 - 0002 - 4827 - 7290 & Paleontological Laboratory, Cherepovets State University, 162600 Cherepovets, Russia; Author Perkovsky, Evgeny E. 0000-0002-7959-4379 Natural History Museum of Denmark, 2100 Copenhagen, Denmark. https: // orcid. org / 0000 - 0002 - 7959 - 4379 * Corresponding author. E-mail: zoya-fedotova @ mail. ru text Ecologica Montenegrina 2024 2024-04-11 72 146 157 http://dx.doi.org/10.37828/em.2024.72.14 journal article 10.37828/em.2024.72.14 2336-9744 13249067 urn:lsid:zoobank.org:pub:63F1C2C7-7296-4CDE-9E53-D83174DB295F Genus Vladimiretskia Fedotova & Perkovsky , gen. nov. https://zoobank.org/ urn:lsid:zoobank.org:act: 4D5A0BD7-5F2D-4D9D-8307-A97EEF6CBCC4 Type species : Vladimiretskia nathani Fedotova et Perkovsky , sp. nov. Diagnosis. Head of female distinctly transverse laterally and frontally. Width of the occipital part of the fused eyes equals 16 facets. Рalpi more than 2.0 times shorter than head height, almost 2.5 times shorter than head length. Antennae 2+16-segmented, almost all flagellomeres with distinct short neck and long basal node. M 1+2 reduced. All legs with femora longer than tibia. First tarsomere with lateral spiniform projection (fig. 2 c, 3 f); tarsal claws on all legs with basal dent (fig. 2 f), dorsally appearing seti-like (fig. 2 e). Lamellae of cerci wide, covered by strongly prolonged setae; apical lamella with 2–3 transparent spines. Description (figs 1–3). Female. Body slightly longer ( 3.2 mm ) than wings ( 3 mm ). Antennae ( 1.26 mm ) more than twice shorter than body plus wing. Head prolonged laterally; eyes very large, occupying almost all lateral side of head, wide divided occipitally, covered basally by long erect setae. Occiput glabrous, occipital setae absent. Clypeus with some erect setae. Mouthparts very short. Palpi 3- segmented, 3 rd segments longest (figs 1 c, 2 a). Antennae 2+16-segmented, scape enlarged distally, pedicel also slightly prolonged. Basal node of 1 st and 2 nd flagellomeres slightly more prolonged than others. All flagellomeres with distinct short, narrow necks, but basal node slightly enlarged distally. Basal nodes of proximal flagellomeres with narrowing near middle, enlarged basally. Medial flagellomeres with very long setae in basal whorls; longest rare setae of medial whorls basally curved. Flagellomeres with circumfilar sensoria in view of basal, apicacal rings, connected by prolonged sensoria. Notum slightly swollen; thorax width, length almost equal. Scutum black-brown, slightly swollen, covered by erect setae. Wing evenly, very strongly widened medially. C with break beyond joining R 5. R 1 joining near mid C . Vein r-m slightly curved medially, R 5 straight apically, joining with margin of wing little before apex; rs undeveloped; M 4 and CuA 1 simple, positioned far from margin of wing, reaching of wing edge; CuA forms large anal lobe; CuP slightly developed. Legs longer than body plus wings, completely, densely covered by short setae and rare long setae. Fore, middle and hind femora longer than their tibia. Ratios of fore tarsi: 1: 9.5: 4.8: 3: 1.5. Tarsal claws with basal dent, slightly curved, with rudimentary empodium. Abdomen strongly swollen basally (1 st –3 rd segments), narrowed apically. Abdominal segments not covered by distinct setae. Tergites, sternites not swollen. Ovipositor very long and thin, VIII–IX segments curved dorsally. Cerci with pair of 3-segmented apical lamellae. Apical lamella of cerci wide, covered by strongly prolonged setae and 2–3 transparent spines (lateral view, figs 1 e, g; 3 j). Etymology . The genus is named for the former Vladimirets District where the amber was mined.