A revision of the genus Belciana Walker, 1862 with description of three new species (Lepidoptera, Noctuidae: Pantheinae) from East and South East Asia. Revision of Pantheinae, contribution XII Author Behounek, G. Author Han, H. L. Author Kononenko, V. S. text Zootaxa 2015 4027 3 341 365 journal article 10.11646/zootaxa.4027.3.2 89c5bda8-87ad-4f90-a154-2de036e9c71e 1175-5326 241473 8B613537-E7D0-4BDB-B168-943C5B54B0BD Belciana hemodoides Holloway, 2009 ( Figs 5, 6 , 33 , 44 ) Belciana hemodoides Holloway, 2009 , The Moths of Borneo 13: 20, Pl. 1: 11, Holotype : male, Brunei , Ulu Temburong, 300 m , rainforest (NHM). References . Kononenko & Pinratana 2005 : 134, Pl. 39:6 ( Belciana biformis ); 2013: 277, Pl. 37: 17 ( Belciana hemodoides ). Material examined. VIETNAM : 2 males , Prov. Lao Cai, Tonkin , Ba Be Lake-National Park, local collector leg., 4.xii.2008 , slides GB12175, GB12176 males, coll. AB; THAILAND : 1 female , 2 females , Prov. Nakhon Ratchasima, Taplan Nationalpark, Bansanchao Po, 520 m , 14°22′544 N, 101°51′ 893 E, Ihle T. leg., 5 -10.v.2005 , slide, GB12186 female, coll. AB; 1 male , 1 female , Prov. Chumphon, Bi Patho, SW Lang Suan, 190 m , 09°46′ N , 98°46′ E , Ihle T. leg., 4.xii.2007 , slide, GB12185 male, coll. AB; 1 female , same place, same collector, 4.xi.2007 , coll. GB/ ZSM ; MALAYSIA , BORNEO : 1 female , Prov. Sabah, Ranau Umg., Poring Hot Spring, Poring Lodge, 06°02′ N 116°42′ E , Chen M.Y. & Buchsbaum U. leg., 7.i.2005 , slide GB7359 female, coll. GB/ ZSM ; PHILIPPINES : LEYTE , 1 male , Mt. Bolocawe, Hilusig, W Mahaplag, 600 m , 10°43′ N , 124°55′ E , Lourens J. H. leg., 19-20.v.2005 , slide, GB/ ZSM N4288 male, coll. ZSM ; LUZON , 1 female , Sierra Madre, Isabela, Dibulo, 13- 17 km N, 550-585 m , 16°32′ N , 122°14′ E , Lourens J. H. leg., 21-22.ix.2006 , coll. ZSM ; 1 female , Sierra Madre, Nueva Ecija boundary, Bongabong Brgy. Laby, 600 m , 15°39′ N , 121°16′ E , Lourens J. H. leg., 5.vi.2005 , coll. ZSM ; 1 male , Sierra Madre, Mingan Mts., Bongabon N. Ec Bgy Laby, 950 m , 12°38′ N , 121°15′ E , Lourens J.H. leg., 7-9.ix.2005 , coll. ZSM ; 1 male , Sierra Madre, Nueva Ecija, Mingan Mts., Bgy. Laby, 600-950 m , 15°38′ N , 121°15′ E , Lourens J. H. leg., 7-9.ix.2005 , slide, GB/ ZSM N4287 male, coll. ZSM ; 1 female , CAR border Abra/ Kalinga, E Malibcong Basiwang, 1550-1700 m , 17°30′200 N, 120°58′881 E, Lourens J. H. leg., 3-4.iv.2008 , slide, GB/ ZSM N4286 female, coll. ZSM ; 1 male , SE of Pagudpud Adams Masasabug, Bgy Bucurat, 400 m , 18°25′769′ N, 120°55′ 259′ E, Lourens J. H. leg., 20.iv.2007 , slide, GB/ ZSM N4285 male, coll. ZSM ; MINDORO , 1 male , Halcon Mts., Mt. Puerto Gallero, Malasembo, 12°22′ N , 121°05′ E , Brechlin R. ex coll., viii.1998 , slide, GB7380 male, coll. GB/ ZSM ; PALAWAN , NE , 1 male , 1 female , Bagong Bayan, foot of Mt. Ilian, 300 m , 10°26′ N , 119°33′ E , Lourens J. H. leg., 8.iii.2006 , coll. ZSM ; 1 male ; Mt. Matalingajan, Napsan, 7 km N Salakot Falls, 950 m , 09°51′ N , 118°37′ E . Lourens J.H. leg., 14-15.iii.2006 , coll. ZSM ; 1 male , Vincente, 20 km ONE Roxas, (primary wood), 400 m , 110°10′N , 10°21′E , Cerny K. & Schintlmeister A. leg., 12-17.i.1988 , slide, GB7389 male, coll. GB/ ZSM ; 1 female , SAMAR , Bagacay, 4 km SE, 140 m , Prim. Forest road, 11°48′ N , 125°15′ E , Lourens J. H. leg., 25.vi.2006 , coll. ZSM . Note. The specimen labelled as BC ZSM Lep 65174 has been used for barcoding. Molecular distance based on the Kimura two-parameter model for COI DNA barcodes between nearest B. hemodoides and B. hreblayi is 5,42%. Diagnosis. Adult ( Fig. 5, 6 ). Wingspan 36–43 mm . The species is close to B. biformis having deeper bluishgreen ground colour of forewing and more developed dark-brown elements of wing pattern in subbasal and subterminal fields. Medial field with pale greyish medial shadow bordered with diffused white lines; orbicular as white diffused spot, reniform as transversal blackish streak, separated for two dots; subterminal line blackish with irregular white bordering, sometimes broken for large separate dashes; apical, subapical and tornal areas brown with blackish dashes, more developed compared with B. biformis . The hindwing are dull greyish-brown, paler yellowish-grey towards base and along anal margin; in some specimens medial discal spot and fascia are traceable. Male genitalia ( Fig. 33 ). Differs from B. biformis by more solid uncus, somewhat narrow and less membranous valva with less developed basal coremata, by stronger ventral margin; wider and more acute shape of harpe. Aedeagus with dorsal branch of carina bearing short finger-like extension (absent in B. biformis ); vesica basally bulbous, tubular apically, with 6–8 moderate cornuti; ventral branch of carina with patch of small spines more dispersed than in B. biformis . Female genitalia ( Fig. 44 ). In the female genitalia antrum narrower than in B. biformis , split; ductus bursae shorter than in B. biformis , sclerotised patch in joining of ductus and corpus bursae more clearly formed and more sclerotised than in B. biformis , having quadrangular shape; the main part of corpus bursae is longer than in B. biformis , with broad clear caudal-lateral extension; the bottom of corpus bursae is slightly taper, rather than be rounded like in B. biformis . Distribution and bionomics. * Vietnam (Lao Cai), * Thailand (Prov. Nakhon Ratchasima, Chumphon), Singapore , Malaysia , Borneo (Prov. Kalimantan, Timur, Prov. Daerah Tingkat and Kalimantan Barat), Brunei (Seria, Ulu Temburong), Indonesia (Great Sunda Islands), * Philippines (Isl. Leyte, Luzon, Mindoro, Palawan, Samar). The species occurs in lowland and mountain areas at elevation range 300–1700 m . Moths flying in January, April, September, December. The most widespread species of the genus, reported here for Vietnam , Thailand and Philippines for the first time.