The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae Author Beaver, Roger A. 161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand. Author Smith, Sarah M. 0000-0002-5173-3736 Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736 smith462@msu.edu text Zootaxa 2022 2022-08-08 5174 1 1 24 journal article 115950 10.11646/zootaxa.5174.1.1 c4bf1061-5af8-4f4a-b4ca-234b6d0ca615 1175-5326 6972960 F80F144B-D1E8-4587-A146-0BACFFE18FB6 Scolytoplatypus raja Blandford Scolytoplatypus raja Blandford, 1893: 440 . Recorded from Bhutan by Schedl (1975 as Scolytoplatypus mikado Blandford ). Schedl made frequent errors in identifying and synonymising species of Scolytoplatypus ( Beaver & Gebhardt 2006 ) . The species, S. mikado , has a more northerly distribution than S. raja , and records of the former species from Bhutan , India and Southeast Asia almost certainly all refer to S. raja ( Beaver & Gebhardt 2006 ) . Distribution. Bhutan , China ( Xizang , Yunnan ), India ( Assam , Himachal Pradesh , Uttar Pradesh , West Bengal ), Laos , Kashmir, Nepal , Pakistan , Taiwan , Thailand , Vietnam , West Malaysia . Biology. The biology and life history of Scolytoplatypus species are described by Beeson (1961) , Browne (1961 a), Beaver & Gebhardt (2006) and Beaver & Liu (2018) . Scolytoplatypus raja is a polyphagous species attacking both coniferous and angiosperm trees ( Beeson 1961 , Maiti & Saha 2009 , Wood & Bright 1992 ).