The Influence of Landscape Heterogeneity - Ground Beetles (Coleoptera: Carabidae) in Fthiotida, Central Greece Author Chapman, Anna Nicola text Biodiversity Data Journal 2014 2 1082 1082 http://dx.doi.org/10.3897/BDJ.2.e1082 journal article http://dx.doi.org/10.3897/BDJ.2.e1082 1314-2828-2-1082 Harpalus (Harpalus) atratus (Latreille, 1804) Distribution Europe (except the north) and the Balkans, where it prefers foothills to alpine regions ( Arndt et al. 2011 ). Notes It is a mesoxerophilous and polyphagous species, which prefers to live in forested areas ( Varvara and Apostol 2008 ). It feeds on a mixture of seeds from agricultural crops, weeds and shrub species ( Lundgren 2009 ). In this study, it was only found in the wheat field in the homogeneous area (n = 1).