<document id="C3559D2808E017FAF933C12D00A6B2AF" ID-CLB-Dataset="4031" ID-DOI="10.5852/ejt.2021.750.1365" ID-GBIF-Dataset="818d0bc4-31a6-4319-86a3-6e6b16ab72df" ID-ISSN="2118-9773" ID-Zenodo-Dep="5451747" ID-ZooBank="CE5D8517-AC04-4AF9-BD78-A2D6E21D33A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630516667691" checkinUser="felipe" docAuthor="Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z. &amp; Ahrens, Dirk" docDate="2021" docId="038B976CFF8DFFE9CA2FCD1C107DCD54" docLanguage="en" docName="ejt-750_94-123.pdf" docOrigin="European Journal of Taxonomy 750 (750)" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1365" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astaena excisipes L. W. Saylor 1947" docType="treatment" docVersion="9" lastPageNumber="101" masterDocId="FFB2EF14FF8BFFEEC84EC925155FCE04" masterDocTitle="Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor" masterLastPageNumber="123" masterPageNumber="94" pageNumber="100" updateTime="1699290068827" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="314924CBACC108D863B000588E9B362C">Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor</mods:title>
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<mods:namePart id="E37E9116AC4C39C2208A73E36E77F176">Pacheco, Thaynara L.</mods:namePart>
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<mods:namePart id="B211D30738D1D040905E788FB940AE1B">Monné, Marcela L.</mods:namePart>
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<mods:affiliation id="565E201314F0C137477F1B4F0C7DAA79">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
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<mods:affiliation id="D337CD8D200D52945E2C22B784EC04DE">Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="53868D445E2544245577A742528368D3" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:affiliation id="15259D087E4214C042E6F233845A45BD">Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
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<taxonomicName id="4C225DF9FF8DFFE8CA2FCD1C168DCA57" authority="Saylor, 1947" authorityName="L. W. Saylor" authorityYear="1947" box="[609,978,1081,1107]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="excisipes">
<emphasis id="B956FA68FF8DFFE8CA2FCD1C1669CA57" bold="true" box="[609,822,1081,1107]" italics="true" pageId="6" pageNumber="100">Astaena excisipes</emphasis>
Saylor, 1947
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<figureCitation id="13193AFFFF8DFFE8CA8ECD79161ACA73" box="[704,837,1116,1143]" captionStart="Fig" captionStartId="8.[189,232,1857,1883]" captionTargetBox="[244,1343,265,1819]" captionTargetId="figure-11@8.[244,1343,265,1819]" captionTargetPageId="8" captionText="Fig. 2. A–D. Astaena excisipes Saylor, 1947, holotype, ♀ (Type No. 7910; CAS). – E–I. A. explaniceps Saylor, 1947, holotype, ♂ (Type No. 7911; CAS). – J–M. A. fusagona Saylor, 1946, holotype, ♀ (Type No. 7912; CAS). A, J. Head and pronotum, dorsal view. B, G, K. labels. C, H, L. Habitus, dorsal view. D, I, M. Habitus, lateral view. E. Parameres, dorsal view. F. Aedeagus, lateral view. Scale bars: A, E–F, J = 0.5 mm; C–D, H–I, L–M = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451751" httpUri="https://zenodo.org/record/5451751/files/figure.png" pageId="6" pageNumber="100">Figs 2A–D</figureCitation>
, 
<figureCitation id="13193AFFFF8DFFE8CB1FCD78162BCA73" box="[849,884,1117,1143]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[261,1326,295,1750]" captionTargetId="figure-12@26.[241,1347,265,1784]" captionTargetPageId="26" captionText="Fig. 7. Distribution of type localities of the South American Sericini species described by Saylor. A. Astaena abaca Saylor, 1946, A. biciliata Saylor, 1946, A. bogotana Saylor, 1946, A. excisipes Saylor, 1947, A. explaniceps Saylor, 1947, A. incachaca Saylor, 1946, A. maqueta Saylor, 1947, and A. nigrona Saylor, 1947. B. A. fusagona Saylor, 1946, A. pygidia Saylor, 1946, A. rugithorax Saylor, 1946, A. salta Saylor, 1946, A. yungasa Saylor, 1946, and A. zyrota Saylor, 1946. C. Sayloria abcora (Saylor, 1946) comb. nov., S. bicoloripes (Saylor, 1946) comb. nov., and S. pottsi (Saylor, 1946) comb. nov.." figureDoi="http://doi.org/10.5281/zenodo.5451761" httpUri="https://zenodo.org/record/5451761/files/figure.png" pageId="6" pageNumber="100">7A</figureCitation>
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<treatmentCitation id="0A83006BFF8DFFE8C8F3CDBA1728CABD" author="Saylor L. W." box="[189,631,1182,1209]" page="433" pageId="6" pageNumber="100" year="1947">
<taxonomicName id="4C225DF9FF8DFFE8C8F3CDBA1728CABD" authority="Saylor, 1947 b: 433" authorityName="L. W. Saylor" authorityPageNumber="433" authorityYear="1947" box="[189,631,1182,1209]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="excisipes">
<emphasis id="B956FA68FF8DFFE8C8F3CDBA14D2CABC" box="[189,397,1183,1209]" italics="true" pageId="6" pageNumber="100">Astaena excisipes</emphasis>
<bibRefCitation id="EFB35B8BFF8DFFE8C9DACDBB1728CABD" author="Saylor L. W." box="[404,631,1182,1209]" pageId="6" pageNumber="100" pagination="433 - 437" refId="ref14000" refString="Saylor L. W. 1947 b. New South American scarab beetles of the genus Astaena. Revista de Entomologia 18: 433 - 437." type="journal article" year="1947">Saylor, 1947b: 433</bibRefCitation>
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.
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<treatmentCitationGroup id="AB320154FF8DFFE8C8F3CDC416ADCB1B" pageId="6" pageNumber="100">
<taxonomicName id="4C225DF9FF8DFFE8C8F3CDC414CFCAFE" authorityName="L. W. Saylor" authorityYear="1947" box="[189,400,1249,1275]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="excisipes">
<emphasis id="B956FA68FF8DFFE8C8F3CDC414CFCAFE" box="[189,400,1249,1275]" italics="true" pageId="6" pageNumber="100">Astaena excisipes</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0A83006BFF8DFFE8C9FDCDC41733CAFE" author="Frey G." box="[435,620,1248,1275]" page="347" pageId="6" pageNumber="100" year="1973">
<bibRefCitation id="EFB35B8BFF8DFFE8C9FDCDC41733CAFE" author="Frey G." box="[435,620,1248,1275]" pageId="6" pageNumber="100" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey 1973: 347</bibRefCitation>
</treatmentCitation>
(identification key). — 
<treatmentCitation id="0A83006BFF8DFFE8CBDCCDC41111CAFE" author="Evans A. V." box="[914,1102,1248,1275]" page="22" pageId="6" pageNumber="100" year="2003">
<bibRefCitation id="EFB35B8BFF8DFFE8CBDCCDC41111CAFE" author="Evans A. V." box="[914,1102,1248,1275]" pageId="6" pageNumber="100" pagination="1 - 458" refId="ref13003" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 22</bibRefCitation>
</treatmentCitation>
(checklist). — Evans &amp; 
<treatmentCitation id="0A83006BFF8DFFE8C8A4CC2114CECB1B" author="Smith A. B. T. &amp; Evans A. V." box="[234,401,1284,1311]" page="7" pageId="6" pageNumber="100" year="2005">
<bibRefCitation id="EFB35B8BFF8DFFE8C8A4CC2114CECB1B" author="Smith A. B. T. &amp; Evans A. V." box="[234,401,1284,1311]" pageId="6" pageNumber="100" pagination="29 - 60" refId="ref14061" refString="Smith A. B. T. &amp; Evans A. V. 2005. A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification. Zootaxa 1032: 29 - 60. https: // doi. org / 10.11646 / zootaxa. 1032.1.2" type="journal article" year="2005">Smith 2005: 7</bibRefCitation>
</treatmentCitation>
(checklist); 2007: 7 (checklist); 2009: 7 (checklist).
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</paragraph>
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<emphasis id="B956FA68FF8DFFE8C8F3CC6314B3CB64" bold="true" box="[189,492,1350,1376]" pageId="6" pageNumber="100">Type material examined</emphasis>
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</paragraph>
<paragraph id="8B9D267AFF8DFFE8C8A4CC50102ACBD2" blockId="6.[189,1399,1397,1495]" pageId="6" pageNumber="100">
<materialsCitation id="3B4A2C27FF8DFFE8C8A4CC50102ACBD2" ID-GBIF-Occurrence="3383726317" collectionCode="CAS" collectorName="Coll. Kraatz" country="Ecuador" determinerName="Moser" location="ECUADOR" pageId="6" pageNumber="100" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B956FA68FF8DFFE8C8A4CC501403CB8B" bold="true" box="[234,348,1397,1423]" pageId="6" pageNumber="100">
<typeStatus id="549998D8FF8DFFE8C8A4CC501403CB8B" box="[234,348,1397,1423]" pageId="6" pageNumber="100" type="holotype">Holotype</typeStatus>
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<collectingCountry id="F33566EAFF8DFFE8C8F3CCBC1414CBB7" box="[189,331,1433,1459]" name="Ecuador" pageId="6" pageNumber="100">ECUADOR</collectingCountry>
• 
<specimenCount id="9D24EDF3FF8DFFE8C910CCBC142ACBB7" box="[350,373,1433,1459]" pageId="6" pageNumber="100" type="female">♀</specimenCount>
; “Ecuador Baron / 
<collectorName id="26D743ACFF8DFFE8CA02CCBD1786CBB7" box="[588,729,1432,1459]" pageId="6" pageNumber="100">Coll. Kraatz</collectorName>
/ L.W. Saylor Collection / California Academy of Sciences Type nº 7910 / 
<taxonomicName id="4C225DF9FF8DFFE8C921CC9817D7CBD2" authority="Bates" authorityName="Bates" authorityYear="1891" box="[367,648,1468,1495]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="B956FA68FF8DFFE8C921CC981761CBD2" box="[367,574,1468,1495]" italics="true" pageId="6" pageNumber="100">Astaena producta</emphasis>
Bates
</taxonomicName>
? / Moser det. / Holotype 
<taxonomicName id="4C225DF9FF8DFFE8CBFDCC981041CBD2" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1947" box="[947,1310,1468,1495]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="excisipes">
<emphasis id="B956FA68FF8DFFE8CBFDCC9811DCCBD2" box="[947,1155,1469,1495]" italics="true" pageId="6" pageNumber="100">Astaena excisipes</emphasis>
L.W. Saylor
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”; 
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.
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</paragraph>
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<typeStatus id="549998D8FF8DFFE8C8F3CCDB1470C81C" box="[189,303,1534,1560]" pageId="6" pageNumber="100" type="holotype">Holotype</typeStatus>
redescription
</emphasis>
</heading>
</paragraph>
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<paragraph id="8B9D267AFF8DFFE8C8F3CF081658C84C" blockId="6.[189,775,1581,1608]" box="[189,775,1581,1608]" pageId="6" pageNumber="100">
MEASUREMENTS. Length: 
<quantity id="4CDA8B9FFF8DFFE8C9A8CF08171CC84C" box="[486,579,1581,1608]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="100" unit="mm" value="9.5">9.5 mm</quantity>
. Width: 
<quantity id="4CDA8B9FFF8DFFE8CAEBCF08165DC84C" box="[677,770,1581,1608]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="100" unit="mm" value="4.7">4.7 mm</quantity>
.
</paragraph>
<paragraph id="8B9D267AFF8DFFE8C8F3CF4A1607C8D5" blockId="6.[187,1399,1647,1745]" pageId="6" pageNumber="100">HEAD. Reddish-brown. Labroclypeus, shape of anterior margin weakly sinuated medially. Ocular canthus with terminal setae, ratio to eye: 1/4. Ratio third vs fourth antennomere: 1/4. Fourth antennomere, lateral projection absent. Ratio fifth vs fourth antennomere: 1/5.</paragraph>
<paragraph id="8B9D267AFF8DFFE8C8F3CFDD1701C9EC" blockId="6.[189,1400,1784,2024]" pageId="6" pageNumber="100">PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, reddishbrown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin straight and not serrated, two angles on postero-lateral margin present, antero-lateral angle dorsoventrally not flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddishbrown, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal.</paragraph>
<paragraph id="8B9D267AFF8CFFE9C8F3C82C17BECF4C" blockId="7.[189,1398,265,329]" pageId="7" pageNumber="101">ABDOMEN. Lateral carina present. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite.</paragraph>
<paragraph id="8B9D267AFF8CFFE9C8F3C85414B4CC6D" blockId="7.[188,1399,369,617]" pageId="7" pageNumber="101">LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, dorsal margin not carinated, ventral margin serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1/2. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid.</paragraph>
<paragraph id="8B9D267AFF8CFFE9C8F3CBB615A2CCA9" blockId="7.[189,253,659,685]" box="[189,253,659,685]" pageId="7" pageNumber="101">
<heading id="D0D59116FF8CFFE9C8F3CBB615A2CCA9" bold="true" box="[189,253,659,685]" fontSize="11" level="3" pageId="7" pageNumber="101" reason="3">
<emphasis id="B956FA68FF8CFFE9C8F3CBB615A2CCA9" bold="true" box="[189,253,659,685]" pageId="7" pageNumber="101">Male</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF8CFFE9C8F3CBE61466CCD9" blockId="7.[189,313,707,733]" box="[189,313,707,733]" pageId="7" pageNumber="101">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF8CFFE9C8F3CA23107DCD54" box="[189,1314,774,849]" pageId="7" pageNumber="101" type="discussion">
<paragraph id="8B9D267AFF8CFFE9C8F3CA231470CD24" blockId="7.[189,303,774,800]" box="[189,303,774,800]" pageId="7" pageNumber="101">
<heading id="D0D59116FF8CFFE9C8F3CA231470CD24" bold="true" box="[189,303,774,800]" fontSize="11" level="3" pageId="7" pageNumber="101" reason="3">
<emphasis id="B956FA68FF8CFFE9C8F3CA231470CD24" bold="true" box="[189,303,774,800]" pageId="7" pageNumber="101">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF8CFFE9C8F3CA13107DCD54" blockId="7.[189,1314,822,849]" box="[189,1314,822,849]" pageId="7" pageNumber="101">
The 
<typeStatus id="549998D8FF8CFFE9C8BCCA131406CD54" box="[242,345,822,848]" pageId="7" pageNumber="101" type="holotype">holotype</typeStatus>
specimen is a female; 
<bibRefCitation id="EFB35B8BFF8CFFE9CA2BCA131659CD55" author="Saylor L. W." box="[613,774,822,849]" pageId="7" pageNumber="101" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor (1946)</bibRefCitation>
incorrectly stated that the 
<typeStatus id="549998D8FF8CFFE9CC71CA1311F9CD54" box="[1087,1190,822,848]" pageId="7" pageNumber="101" type="holotype">holotype</typeStatus>
as a male.
</paragraph>
</subSubSection>
</treatment>
</document>