The earliest beetle with mouthparts specialized for feeding on nectar is a parasitoid of mid-Cretaceous Hymenoptera Author Batelka, Jan Department of Zoology, Faculty of Science, Charles University, ViniČná 7, 128 00 Praha 2, Czech Republic janbat@centrum.cz Author Prokop, Jakub Department of Zoology, Faculty of Science, Charles University, ViniČná 7, 128 00 Praha 2, Czech Republic text BMC Ecology and Evolution 2021 207 2021-11-22 21 1 1 12 http://dx.doi.org/10.1186/s12862-021-01930-6 journal article 10.1186/s12862-021-01930-6 b2e4e166-7297-45a5-82eb-84937eb7233a PMC8607574 34809578 5733385 Melanosiagon gen. nov. Type species. Melanosiagonserraticornis sp. nov. Etymology . Generic name is a combination of the black appearance of the specimen (Greek μελάνω) and the generic name Macrosiagon [ 28 ]. Feminine in gender. Te name is registeredunder ZooBank LSID urn:lsid:zoobank. org:act:46B9D135-5214-4999-9F2B-0BF4133F5F7F. Diagnosis. Tis genus is characterized by having a long and slender body, long elytra, acute, convex and partially dehiscent, small head, strongly flattened antero-posteriorly with vertex transversely planar, specialized mouthparts with elongated galea, bilobed ligula covered by bristles, prominent tibial spurs, tibial spur formula 2-2-2, antennae 11-segmented, mesonotal and metanotal legs with erect stiff spiniform setae, pretarsal claws serrate, abdominal segments laterally compressed, apical fields on hind wings with secondary “ghost” branches (sensu [ 26 ]: 186) and apices of hind wings asymmetrically overlapping in resting position.