<document id="026250AA1321C2089207664C058134AA" ID-CLB-Dataset="302074" ID-DOI="10.3897/phytokeys.246.129476" ID-GBIF-Dataset="b381880a-e0bb-4b02-ad0f-2f90a0b6c995" ID-publisher-id="129476" URI-arpha="7631F16E-4CD8-5A76-9F28-45E5F398CE20" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1725298343909" checkinUser="pensoft" docAuthor="Meerts, Pierre J. &amp; Paton, Alan J." docDate="2024" docId="3DC4DBC3DAB85365B0D61DFFEB1B79CF" docLanguage="en" docName="PhytoKeys 246: 71-178" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/129476/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Coleus kivuensis Lebrun &amp; L. Touss." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="178" masterDocId="7631F16E4CD85A769F2845E5F398CE20" masterDocTitle="The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species" masterLastPageNumber="178" masterPageNumber="71" pageNumber="71" updateTime="1725322260938" updateUser="ExternalLinkService">
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<mods:title id="05AABA19A97270BAE3E3E031ECFD830F">The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species</mods:title>
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<mods:namePart id="C416307ADEBA361A5A2775072534B5A7">Meerts, Pierre J.</mods:namePart>
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<mods:affiliation id="58C110DB50AAD3FDAE4107E893A6CCEF">Meise Botanic Garden, Nieuwelaan 38, BE- 1860 Meise, Belgium &amp; Université Libre de Bruxelles, Av. F. D. Roosevelt 50 CP 244, BE- 1050 Brussels, Belgium &amp; Fédération Wallonie-Bruxelles, Service général de l’Enseignement supérieur et de la Recherche scientifique, Brussels, Belgium</mods:affiliation>
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<mods:namePart id="5123828458522132321A01CF0A26BA3E">Paton, Alan J.</mods:namePart>
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<mods:affiliation id="87F5E10354DA37A377085FC5E693F496">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
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<mods:title id="946B6F8C38FF8DEC281300BB0D29DED2">PhytoKeys</mods:title>
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<mods:part id="F800378BAA6A449B95AF567FD86D1564">
<mods:date id="1F300ABB92841B10B8DFB3F7AA9D7E6F">2024</mods:date>
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<mods:classification id="C5F6484A18931195C54D9D04E1E25955">journal article</mods:classification>
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<mods:identifier id="0CEEDFFD7487E3AFD22E6F5B12924936" type="DOI">10.3897/phytokeys.246.129476</mods:identifier>
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<treatment id="3DC4DBC3DAB85365B0D61DFFEB1B79CF" ID-DOI="http://doi.org/10.5281/zenodo.13629821" ID-GBIF-Taxon="240061455" ID-Zenodo-Dep="13629821" ID-arpha="3DC4DBC3-DAB8-5365-B0D6-1DFFEB1B79CF" LSID="urn:lsid:plazi:treatment:3DC4DBC3DAB85365B0D61DFFEB1B79CF" httpUri="http://treatment.plazi.org/id/3DC4DBC3DAB85365B0D61DFFEB1B79CF">
<subSubSection id="21711F6EDC673DDEDBC9FBE22C806BA6" type="nomenclature">
<paragraph id="FF7DE5C26BA46FEDED5AC354DF2465BC">
<treatmentCitation id="E70AD97155919CD269A18216560FAAF4" author="Lebrun &amp; L. Touss." journal="Bull. Jard. Bot. Etat Bruxelles" page="72" score="291.0324320782438" volume="17" year="1943">
<taxonomicName id="4AC8F5608C83FD11B5EAA4C8635E70BC" ID-CoL="X26X" authority="Lebrun &amp; L. Touss." authorityName="Lebrun &amp; L. Touss." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="kivuensis">
<emphasis id="E415232F0D999257190CCE4DB5B484ED" italics="true">Coleus kivuensis</emphasis>
Lebrun &amp; L. Touss.
</taxonomicName>
, Bull. Jard. Bot. État Bruxelles 17: 72. 1943
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="937599ED631D71E77500397E7C6A3F38" type="reference_group">
<paragraph id="69CBE65D50B9C01F19FB8BE3C3D6F394">
<treatmentCitationGroup id="DA34BED24E4024979C89F8B19E79473F">
≡ 
<treatmentCitation id="6F181F1C4FF7A97511C6A64C4325FC8C" author="R. H. Willemse" journal="Kew Bull." page="96" score="22.239024375395413" volume="40" year="1985">
<taxonomicName id="9C408C9D41977D6B5A2AE6369A2C676C" ID-CoL="4K4CD" authority="(Lebrun &amp; L. Touss.) R. H. Willemse" authorityName="R. H. Willemse" baseAuthorityName="Lebrun &amp; L. Touss." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="kivuensis">
<emphasis id="1D054D2DC8CE9E4B9255458D47315266" italics="true">Plectranthus kivuensis</emphasis>
(Lebrun &amp; L. Touss.) R. H. Willemse
</taxonomicName>
, Kew Bull. 40: 96. 1985
</treatmentCitation>
. Type: 
<collectingCountry id="C771BA878BFBF0A836DFE794B5677D80">DR. Congo</collectingCountry>
, Kivu, Rutshuru, 
<date id="80F512843ED3FD42A1CF1E6AB38DF15E" value="1937-12">Dec 1937</date>
, 
<emphasis id="044458EDD1899D950DAE0E77CCAD30C5" italics="true">J. Lebrun 9031</emphasis>
(
<typeStatus id="555EBCD3CC5A53FB9B3842F8A0D0BB87" type="holotype">holotype</typeStatus>
<collectionCode id="ACD99915FF9AE60072A1A1C230652BBD" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="FBD15621E558315747ADC231AF7147DE">
<collectionCode id="2DAFA23C6B89123FB3C3F7D095EC3A1A" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000006262563
</specimenCode>
]; 
<typeStatus id="E8F8F8B255366E50F2F1A103E3390A99" type="isotype">isotype</typeStatus>
<collectionCode id="F8B63AC4FAB37C575B529BC742F23AE8" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
[
<specimenCode id="F9150D292098E050DFAD21C9AE281C81">
<collectionCode id="45AA0C534EBE01A7EF4474E9169E5D35" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
000431876
</specimenCode>
], [
<specimenCode id="040F953828D8EC89F9CE81F16B66CE9C">
<collectionCode id="CB9C8884676933A7EE0CE951DA82BD7D" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
000431875
</specimenCode>
], P [P 00450798]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2FF42784943D64BB9B1E10D3FF875192">
<treatmentCitationGroup id="A83AE2AC3EEFBA3A39CAAFDCEB69A4B8">
<comment id="E413BBE970A280FBB4112AAAEF6A49A7">
= 
<taxonomicName id="10F83AFB8D29FEEB609473363B29F021" ID-CoL="4K4FV" authority="sensu Troupin &amp; Ayob." authorityName="Troupin &amp; Ayob." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="neochilus">
<emphasis id="30E3208125AFA26E09BDC9AF3792E19D" italics="true">Plectranthus neochilus</emphasis>
sensu Troupin &amp; Ayob.
</taxonomicName>
, 
<treatmentCitation id="FEEDAAFF843CDC9A785BA561F27869B6" author="Troupin &amp; Ayob., Fl." journal="Rwanda" page="339" score="1.97023359760351E-6" volume="3" year="1985">
Fl. 
<bibCitation id="53F4C4B875B1C68A71DE47FD25848A0D" journal="Rwanda" page="339" score="1.97023359760351E-6" volume="3" year="1985">
<collectingCountry id="41A43EDD046D7022562DDCD3476625C2" name="Rwanda">Rwanda</collectingCountry>
3: 339. 1985
</bibCitation>
</treatmentCitation>
., non Schltr.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERIBM" type="description">
<paragraph id="DF231317EB1F7862BA9774CD2CCADC24">
<heading id="8701FF9F030398838D6E043A16372107" reason="title">Description.</heading>
</paragraph>
<paragraph id="21D63BAB9A5A5DB9A9DBA79A105AF0BF">
<bibRefCitation id="B5E017A78F927EA1A9CFCE5FD9F21719" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Beentje HJ &amp; Ghazanfar SA &amp; Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="346" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1–430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009: 346)</bibRefCitation>
, as 
<taxonomicName id="4DE43BAA6D276EC3F2C6A0858B34B277" authority="(Lebrun &amp; L. Touss.) Willemse." authorityName="Willemse." baseAuthorityName="Lebrun &amp; L. Touss." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="kivuensis">
<emphasis id="6B999338578F158A6C57EB240E1E1431" italics="true">Plectranthus kivuensis</emphasis>
(Lebrun &amp; L. Touss.) Willemse.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGJBM" type="distribution">
<paragraph id="EA72CF411227CEC618D2A1A4AE9469F3">
<heading id="AB3036F9082913B944C3D03554C76723" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="ECF1D8AB5336732C9360DCCD6CC0C43A">
<collectingCountry id="EC696FDAD6F57BA260752DABBF9F06F7" name="Eritrea">Eritrea</collectingCountry>
to N. 
<collectingCountry id="90FB75F115B91F56926670E958153FCE" name="Tanzania">Tanzania</collectingCountry>
and 
<collectionCode id="16A90670FD16E62EFC7818D2AE8B1D24" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
<collectingCountry id="95049C6B64690CB54CB9D2DEAFD8510B">DR. Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESJBM" type="habitat">
<paragraph id="068D252CB7F6A2243EFFAD7D90398942">
<heading id="07A49DD94906CE9405EDBDD7C94DA573" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="D64E13EF511A817C0625BC71D927AF4D">Shrub savannah, fallow fields, steppe; 900–1400 (– 2200) m elev.</paragraph>
</subSubSection>
<subSubSection id="SECID0EXJBM" type="Additional specimens">
<paragraph id="7090A17C0A921AACF18AA352A9816EB8">
<heading id="8023CF1F7AFFF3877C7B19B3DAC2011F" reason="title">Additional specimens.</heading>
</paragraph>
<paragraph id="7880BF088A44E141B44226B725680795">
<collectingCountry id="DD5D0002364A1F0E1EBB00F84C8E706B">DR. Congo</collectingCountry>
, 
<emphasis id="8E24BBE560E9A637F28876C921002AB1" bold="true" italics="true">Lacs Edouard et Kivu</emphasis>
, Kengele, pied du Ruwenzori, 
<date id="A8AB39126DB0024634A670C94D3E0A44" value="1914-04-27">27 Apr 1914</date>
, 
<emphasis id="F6CEAB0E04A5985BBC89A8293ADA2153" italics="true">J. Bequaert 3969</emphasis>
(
<collectionCode id="89FB94D4BC91748BBA78268F74DAA5F0" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Keshero, 
<date id="76DD95D8688513C1E584C011E814094E" value="1958-09-23">23 Sep 1958</date>
, 
<emphasis id="B47FE80DBED24F1C42A9831FBC472985" italics="true">Crispiels-Thonon 124</emphasis>
(
<collectionCode id="023020111560C8F8FDDA85FF6771908D" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Près de Kambukabakali, rive droite de la Semliki, 
<date id="594A104F5F67CEFB6F782510EEB0F430" value="1954-10-27">27 Oct 1954</date>
, 
<emphasis id="F330D4CAAD271B15EE80FA8D61933147" italics="true">
<collectionCode id="A4189CAF0F8C2725CC6B1B0F496CBF43" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. F. de Witte 11278
</emphasis>
(
<collectionCode id="57F269F3E8E55122D3ED13D8D4E618DE" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Kaliba, pied du Kasali, 
<date id="581A783B5B931449B6138D1542DE8683" value="1957-03-11">11 Mar 1957</date>
, 
<emphasis id="6A5E2F450B800AD577E4804F1C3348E3" italics="true">
<collectionCode id="D63F8548AB4817C60737C6D7F578E43C" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. F. de Witte 14035
</emphasis>
(
<collectionCode id="6ECCF58C68418FC08090374FD60B4524" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Rutshuru, 
<date id="48173D2BD06C760512FAD5CD4DD35B10" value="1971-11-16">16 Nov 1971</date>
, 
<emphasis id="7B9EFF991F2B649BF99B00EEBC7DF382" italics="true">
<collectionCode id="DF449A72ED56F407A8DF6B474A6F760E" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
. Evrard 6827
</emphasis>
(
<collectionCode id="DC568D4027C5E53394E5228452187CC7" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Rutshuru, 
<date id="25F311F5718C2105A60501F762C31D97" value="1936-12-30">30 Dec 1936</date>
, 
<emphasis id="A69B2516297F5CA27ABD5B74D1A69048" italics="true">J. Ghesquière 3573</emphasis>
(
<collectionCode id="18B95D7B03A51730D745F752DC7B0E10" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
, 
<collectionCode id="1B453C17C44AF09158CF1D2D05BAF384" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
).
</paragraph>
<paragraph id="B40EE81099A8085CE962AA83F9361319">
<collectingCountry id="840F727DC956FD70E723F9589C1BBAFB" name="Rwanda">Rwanda</collectingCountry>
, Mulehe (Bugesera), 
<date id="57A4D5D8CA767EFB272CF8F44F2C60AE" value="1954-03-15">15 Mar 1954</date>
, 
<emphasis id="912F0AB83C158AAC377878B60A2B864F" italics="true">L. Liben 1268</emphasis>
(
<collectionCode id="FF9F8A8E958B6436554694696B230241" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
, 
<collectionCode id="AF3C15988BFDC84DEAFF247CF35D31E6" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
); Parc national de l’Akagera, plaine de Nyaruhuru, 
<date id="5468758FEDF4B7500F8A05A22687543F" value="1969-04-14">14 Apr 1969</date>
, 
<emphasis id="41183F9E7B8315B69484A327DDC8D3BF" italics="true">
<collectionCode id="4FD1007404003C60AE5EAA3DADBE67A6" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Bouxin &amp; 
<collectionCode id="E33CDBBC1CA82C100386EF525474A828" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Radoux 167
</emphasis>
(
<collectionCode id="591A64BA63A6B03F443FA73EBFE28EB6" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Région du Matara, environs de Mimuli, 
<date id="28ED5E3B0B4B19DAA6829016485E69C2" value="1957-05-23">23 May 1957</date>
, 
<emphasis id="92DA7EF6A7011B83316481CB9544D66E" italics="true">
<collectionCode id="754531B7B3F6BE23E9435C7FDF7EEE03" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Troupin 3205
</emphasis>
(
<collectionCode id="BB7055126A47DE6565A4FC6AC57A2266" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Région du Mutara, environs de Nyagatare, colline Rutare, 
<date id="4B89989C2D865941EC925333F71A4614" value="1958-05-01">1 May 1958</date>
, 
<emphasis id="4045AF362EFC4908EAA8FF444F3A4CD4" italics="true">
<collectionCode id="44C5F5EEBEA134F657A42DCD88EF76B5" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Troupin 7239
</emphasis>
(
<collectionCode id="4EFBB800EDC67D05A66D80BF52877A48" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>