Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species
Author
Marzuki, Mohammad Effendi bin
Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88450, Kota Kinabalu, Sabah, Malaysia
fendiemz@gmail.com
Author
Liew, Thor-Seng
https://orcid.org/0000-0002-9437-5924
Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88450, Kota Kinabalu, Sabah, Malaysia
thorsengliew@gmail.com
Author
Mohd-Azlan, Jayasilan
Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia
text
ZooKeys
2021
2021-04-27
1035
1
113
http://dx.doi.org/10.3897/zookeys.1035.60843
journal article
http://dx.doi.org/10.3897/zookeys.1035.60843
1313-2970-1035-1
ED19022EA1704DB79587FEFE15D07854
4C2258D4EE6754488B9280D3AB0447A1
Everettia microrhytida
sp. nov.
Figures 41B
, 42D-F
Material examined.
Holotype
(SH
12.14 mm
, SW 22.00 mm) (MZU.MOL.20.25),
Malaysia
, Sarawak,
Kuching Division
,
Gunung Batu
, limestone outcrop along
Skio
road,
Jambusan
,
2.4 miles
E Bau
,
1°23'50.65"N
,
110°11'19.99"E
, coll.
M. E. Marzuki
,
10.II.2017
.
Paratypes
:
1 ex.
(MZU.MOL.20.26), the same locality as holotype, coll.
M. E. Marzuki
,
10.VII.2011
;
1 ex.
(ME0006829), small limestone escarpment near
Kampung Padang Pan
,
15 miles
SW Bau
,
1°19'24.07"N
,
110°3'46.34"E
, coll.
M. E. Marzuki
,
27.IX.2015
;
2 ex.
(ME0003498), small limestone outcrop at
Kampung Beratok
,
Serian-Kuching
road,
14.3 miles
NW Serian
,
1°18'41.05"N
,
110°24'37.13"E
, coll.
M. E. Marzuki
,
21.VI.2010
;
6 ex.
(ME0009145),
Lobang Angin
(
Site
3), limestone outcrop near
Sungai
Sarawak
Kanan
,
1.75 miles
W of Bau
,
1°24'54.96"N
,
110°8'13.62"E
, coll.
M. E. Marzuki
,
23.IV.2017
;
1 ex.
(ME0009845), the same locality, coll.
M. E. Marzuki
,
12.V.2018
.
Differential diagnosis.
This species is similar to
Everettia consul
(Pfeiffer, 1854) in terms of general shape and size. However, it differs from
E. consul
by lacking spiral sculpture and having only very fine (sometimes inconspicuous), somewhat wrinkled, puncture-like sculpture on both the apical and apertural sides.
Everettia consul
, on the other hand, has a shell with a more elevated spire and its shell surface has densely placed radial threads and somewhat cut by irregularly spaced spiral grooves.
Figure 42.
Everettia
spp.
A-C
Everettia minuta
, sp. nov., MZU.MOL.20.24 Paratype
A
Enlargement of the apical side showing the apex
B
Enlargement of the teleoconch showing the shell sculpture
C
Enlargement of the basal side of the shell
D-F
Everettia microrhytida
, sp. nov., MZU.MOL.20.26 Paratype
D
Enlargement of the apical side showing the apex
E
Enlargement of the teleoconch showing the shell sculpture
F
Enlargement of the basal side of the shell.
Description.
Shell moderately large, rather thin, translucent, pale to dark brown, spire slightly elevated. Surface with a glossy lustre. Whorls convex. Number of whorls <
61/4
. Protoconch: almost smooth, spiral striation absent with inconspicuous radial threads near the suture. Teleoconch with no: spiral sculpture but with very fine, inconspicuous, somewhat wrinkled, puncture-like shell sculptures on both apical and apertural sides. Radial sculpture of teleoconch very fine as well as inconspicuous growth lines, most conspicuous radial threads near the suture and below periphery. Periphery round; suture shallow. Aperture lunulate. Peristome simple; somewhat thickened and reflected on columellar side, not thickened nor reflected on basal and palatal sides. Umbilicus open, narrow; sometimes partly covered by reflected peristome; umbilical region moderately concave. Dimensions: shell height <12.58 mm; shell width <22.13 mm; diameters of the first three whorls 1.60 mm, 2.55 mm, and 4.90 mm, respectively; aperture height <8.94 mm; aperture width <11.66 mm.
Geographic distribution and habitat.
It is known from the Bau and Serian-Padawan limestone hill clusters. Living snails were observed foraging among leaf litter and plant debris near the cliff in a lowland limestone forest.
Etymology.
From the Greek
mikro
rytides
, meaning extremely small wrinkles, in reference to the shell sculpture.