Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species Author Golovatch, Sergei Russian Academy of Sciences, Moscow, Russia Author Geoffroy, Jean-Jacques urn:lsid:zoobank.org:author: Muséum national d’Histoire naturelle, Département Ecologie & Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France & D 362 Dc 0 - & CC Author Mauriès, Jean-Paul urn:lsid:zoobank.org:author: Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France & - & - 42 F & D 01 - 28 Ad- 43 F Author VandenSpiegel, Didier Musée Royal de l’Afrique centrale, Tervuren, Belgium & A & EE & F- 04802 E 68 Cb text ZooKeys 2009 2009-06-18 12 12 1 46 journal article 10.3897/zookeys.12.167 3026c846-a607-4826-b2a9-8f7a531baec5 1313–2970 576460 B32EF8D5-C3C6-4C8C-8845-284125448425 Eutrichodesmus reductus Golovatch, Geoffroy, Mauriès & VandenSpiegel , sp. n. urn:lsid:zoobank.org:act: 1726A727-8FC2-4BF8-807D-8DFC1C54380C Figs 31-33 Type material: Indonesia , Sulawesi Selatan , kab. Maros : Samanggi , Gua Saripa Cave , hand collected, 18.VIII.1990 , leg. A. Bedos and L. Deharveng (SULS-214), holotype ♁ (MZB), paratypes : 1 ♁ (MNHN JC 325), 1 ♀ (SEM) . Name : To emphasize the strongly underdeveloped paraterga 2 and lack of metatergal tuberculation. Diagnosis : Differs from congeners except E. communicans Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2009 by the strongly underdeveloped paraterga 2, coupled Figure 31 . Eutrichodesmus reductus sp. n. , ♁ and ♀ paratypes ; habitus, lateral view. (Photographed not to scale by L. Deharveng). with 19 body segments and the absence of metatergal tuberculation; from E. communicans by a very short tergal trichome, from it and other congeners in a few minor details of gonopod structure (in particular, the shape of the telopodite). Description : Length of adults of both sexes ca 4.0- 4.2 mm , width 0.45-0.5 mm , body broadest at midbody segments. Holotype ca 4.0 mm long and 0.45 mm wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (Fig. 31). Adults with 19 segments, body subcylindrical (Figs 31, 32A), volvation apparently incomplete due to insufficiently wide and low paraterga. Head ( Fig. 33A ) slightly transverse, with a poorly separated pair of very low, paramedian tubercles above antennal sockets; antennae relatively short and clavate, antennomere 6 longest; tegument ( Figs 33 C-E) and many other characters ( Figs 32F , 33B ) much as in E. distinctus sp. n. ; collum and following metaterga devoid of tuberculation, beset with numerous, irregularly arranged, extremely short setae ( Fig. 32 B-F, 33C-E). Paraterga directed ventrolaterad, slightly interrupting contour of convex dorsum, short, not reaching level of venter ( Figs 32 A-D); paraterga 2 ( Fig. 32B ) only a little enlarged compared to following ones, indistinctly lobulate both anterolaterally and laterally, with only a single noticeable lobe forming a schism ledge, both schism and hyposchism very small; paraterga 3 and 4 not narrower than others, overlap typical already from paraterga 4, not paraterga 5 as in preceding congeners. Paraterga broadly rounded caudally, very indistinctly lobulate laterally and with a single evident lobulation caudolaterally ( Figs 32 C-E, 33C). Limbus distinctly denticulate, almost hidden by nearby abundant microvilli ( Fig. 33D, E ). Pore formula normal, ozopores located dorsally near base of caudal corner of paraterga ( Fig. 33C ). Legs relatively long and slender, evidently surpassing edge of paraterga ( Fig. 33B ). Gonopods ( Fig. 33F, G ) relatively complex. Coxae abundantly micropapillate, but only with a few macrosetae. Telopodite elongate, only slightly arcuate, with a large, papillate, peculiar, distofemoral outgrowth (dp) in distal one-third and a slender but short solenomere bearing a few small setae (but no pad!) subapically at base of a very complex tip represented by two erect teeth, a longer uncus and a group of minute outgrowths. Figure 32. Eutrichodesmus reductus sp. n. , ♀ paratype; A , habitus, lateral view; B , anterior part of body, lateral view; C , midbody segments, lateral view; D-F , posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm. Remarks : This small-bodied, pallid, non-volvatory (= “haplodesmid”, see Golovatch et al. 2009 ) species seems to be especially close, morphologically as well as geographically, to E. communicans from Vanuatu , Melanesia, southwestern Pacific ( Golovatch et al. 2009 ), possibly representing still one more troglobite.