Reassessing the phylogeny and divergence times of sloths (Mammalia: Pilosa: Folivora), exploring alternative morphological partitioning and dating models Author Casali, Daniel M Author Boscaini, Alberto Author Gaudin, Timothy J Author Perini, Fernando A text Zoological Journal of the Linnean Society 2022 2022-12-01 196 4 1505 1551 https://academic.oup.com/zoolinnean/article/196/4/1505/6617197 journal article 10.1093/zoolinnean/zlac041 0024-4082 7381236 Megatheriini PP = 100, age = 9.85 Mya (9.07–10.67). This clade was consistently recovered in all analyses performed in this study with an invariable arrangement:( Anisodontherium ,( Pyramiodontherium , ( Megatheriops , ( Megatherium , ( Proeremotherium , Eremotherium ))))). Megatheriini was supported by eight synapomorphies, all but one, recovered with both methods: thickness of cementum much larger than orthodentine; mf3 smaller than next smallest molariform; reflexed basicranial/basifacial axis; profile of nasal region and braincase relatively horizontal, but nasal region depressed relative to braincase; maximum length of nasal bones greater than or equal to three times the width of both nasals (BI); presence of ventral extension in maxilla for dental alveoli; concave mediolateral contour of palate; and occipital condyles situated well dorsal to the dentition.