Reassessing the phylogeny and divergence times of sloths (Mammalia: Pilosa: Folivora), exploring alternative morphological partitioning and dating models Author Casali, Daniel M Author Boscaini, Alberto Author Gaudin, Timothy J Author Perini, Fernando A text Zoological Journal of the Linnean Society 2022 2022-12-01 196 4 1505 1551 https://academic.oup.com/zoolinnean/article/196/4/1505/6617197 journal article 10.1093/zoolinnean/zlac041 0024-4082 7381236 Choloepodini PP = 98, age = 7.07 Mya (5.67–8.93). This clade is composed of the genus Pliomorphus ; the Antillean megalonychines Megalocnus , Parocnus , Neocnus and Acratocnus ; and the living sloth genus Choloepus . It was consistently recovered in our analyses with the arrangement ( Pliomorphus , (( Megalocnus , Parocnus ), ( Neocnus , ( Acratocnus , Choloepus )))). In one exception—in par_EW— Neocnus , Acratocnus and Choloepus were recovered in a polytomy. Choloepodiniwassupportedbyeightsynapomorphies (six for both methods and two exclusively for BI): snout downturned anteroventrally, in lateral view; evenly convex dorsal surface of the skull; minimum width of palate between toothrows equal or less than width of Mf2; absence of postorbital process of zygomatic arch; occiput wider than deep; presence of recessus meatus (BI); presence of a strong concavity between greater trochanter and the head of femur (BI); and patellar trochlea confluent with lateral, but not with medial condylar surface.