Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan Author Bänziger, Hans Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034 Author Sati, Pradeep C. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575 Author Fartyal, Rajendra S. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263 Author Suwito, Awit Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com Author Katoh, Toru Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545 text Zootaxa 2020 2020-06-08 4789 1 1 54 journal article 21673 10.11646/zootaxa.4789.1.1 c12aaed8-264c-459c-b1aa-09eaeb686670 1175-5326 3884610 311A27A2-A053-4D20-A43E-EF57DBE55166 24) Phortica ( Ashima ) foliacea ( Tsacas & Okada, 1983 ) Amiota ( Phortica ) foliacea Tsacas & Okada, 1983: 233 . Phortica foliacea : Chen et al ., 2005b: 420 . Phortica ( Ashima ) foliacea : Chen & Máca, 2012: 507 . Diagnosis. Arch on anterior margin of aedeagal sheath narrow, less sclerotized; postgonites small; posterior one only on either side; anterior ones articulated to tips of arch, slightly curved inward, subequal in size (“Figs 13, 14” in Tsacas & Okada 1983 ). Supplementary description (not repeating characters common to P . semiannulata ). Supracervical setae 15– 16. Longest, dorsal branch of arista as long as longest seta on pedicel; apical, leaf-like expansion triangular, slightly narrower than long. Cibarial, posterior sensilla 7–8 per side. All tarsi with gray tarsomeres IV and V. The antisymmetry is observed in the postgonites: in A-type, the posterior postgonite only on the right side; but in B-type, only on the left side. Specimens examined. Taiwan : 1♂ (A-type) 2♂ (B-type), Shanlinxi , 16.x.1992 , M.J. Toda leg. ( SEHU ) ; 2♂ (A-type) 1♂ (B-type), Chitou , 21–22.iv.1997 , M.J. Toda leg. ( SEHU ) . Distribution. Taiwan . Remarks. This species resembles P . semiannulata in having the anterior, more or less inward oriented postgonites articulated to tips of sclerotized arch on the anterior margin of aedeagal sheath, but can be distinguished from it by the diagnostic characters.