Taxonomic changes in the emerald moths (Lepidoptera: Geometridae, Geometrinae) of East Asia, with notes on the systematics and phylogeny of Hemitheini Author Beljaev, Eugene A. text Zootaxa 2007 1584 55 68 journal article 10.5281/zenodo.178403 fcd42514-18b6-44e8-b14e-173798c335c6 1175-5326 178403 Microloxia chlorissoides ( Prout, 1912 ) , comb. n. Mixochlora chlorissoides Prout, 1912 , in Wytsmann, Gen. Ins. 129: 201. Holotype : ɗ (BMNH), China : Hong Kong , Happy Valley. Chlorissa chlorissoides : Prout, 1934 , in Seitz, Grossschmett. Erde 12: 177. Nipponogelasma chlorissoides : Inoue, 1971 : 145 , pl. 1: 4, pl. 2: 25, 25a, 26; Inoue, 1977 : 235 ; 1982: 436, pl. 59: 17, 18, Cat. 2211; Parsons et al., 1999 : 645 . Taxonomy. This species was examined in detail by Inoue (1971) , who first described and illustrated the male and female genitalia. Based mostly on facies and wing venation, Inoue (1971) tentatively assigned the species to Nipponogelasma . However, the male of Mixochlora chlorissoides has a large conical distal process on the sacculus, which occurs very rarely in Geometrinae . In the Palaearctic region and in South East Asia this process is known only in the genus Microloxia Warren, 1893 . The male genitalia of Mixochlora chlorissoides share with those of Microloxia herbaria (Hübner, 1813) (the type species of the genus) the following characters: uncus with short bifurcation at the apex; socii similar to uncus in length and shape; valva simple, elongate, with long, broad, slightly convex costa; sacculus with long, conical distal process reaching 4/5 length of valva; transtilla broad, platelike; saccus long, conical; and aedeagus simple, lacking cornuti (morphological data on Microloxia from the description and figures in Hausmann, 1995 , 2001 ). In general, the male genitalia of Mixochlora chlorissoides are very similar to those of Microloxia herbaria ; they differ from the latter by the narrower socii, broader saccus, longer basal process of the aedeagus, and longer ventral processes of the transtilla. However, males of Mixochlora chlorissoides possess a long needlelike posterior process on the eighth abdominal sternite that is absent in other species of Microloxia . In the female genitalia, Mixochlora chlorissoides differs from Microloxia herbaria in the possession of a short ductus bursae and a rounded sacklike corpus bursae, but both species share a very characteristic signum consisting of two small, toothed plates. External characters provide support for the transfer of Mixochlora chlorissoides to Microloxia , including Inoue's note on similarity of wing venation of Mixochlora chlorissoides Prout with that of " Nipponogelasma immunis " (= Aoshakuna lucia ). In the last species the forewing has a very short free basal portion of R1 fused with Sc for a long distance, a condition that rarely occurs in Geometrinae ; the same configuration of Sc–R1 is shown on the figure of the wing venaton of Microloxia herbaria and Microloxia simonyi (Rebel, 1894) by Hausmann (1995: figs 1, 2) . Distribution. Japan (Ryukyu), China (Southeast, Hainan), Taiwan , North Vietnam (" Tonkin ").