An annotated catalogue of the Buprestidae of Iran (Coleoptera: Buprestoidea)
Author
Ghahari, Hassan
Author
Volkovitsh, Mark G.
Author
Bellamy, Charles L.
text
Zootaxa
2015
3984
1
1
141
journal article
10.11646/zootaxa.3984.1.1
30f05c9f-072f-440d-8ac5-b07d47a3f387
1175-5326
245157
318FD9EA-E85D-4AEF-8E02-15EE13295F1B
Genus
JULODIS
Eschscholtz, 1829
synonym:
Jalodis
Eschscholtz 1829.
synonym:
Phyllis
Gistel 1834
.
synonym:
Saccosoma
Motschulsky 1859
.
Note.
The larvae of
Julodinae
are soil habitants feeding externally on the roots of different plants. All the records herein refer to adults only.
Julodis aeneipes
Saunders 1869
: 2
;
Semenov-Tian-Shankij 1906
: 153
;
Radjabi 1968
: 72
;
Abai & Adeli 1984
: 116
;
Modarres Awal 1997
: 135
;
Borumand 2002
: 1
;
Kubáň & Volkovitsh 2006
: 326
;
Bellamy 2008a
: 46
;
Holyński 2014
: 20
.
synonym:
afghanica
Obenberger 1924c
: 7.
Host plant(s) within
Iran
.
(Adult!) Tamarisk (
Tamarix
sp.,
Tamaricaceae
) (
Abai & Adeli 1984
;
Modarres Awal 1997
).
Distribution within
Iran
.
Bushehr (
Obenberger 1923a
), Hormozgan, Kerman (
Borumand 2002
), Sistan & Baluchestan (
Semenov-Tian-Shankij 1906
;
Obenberger 1924c
, as
afganica
; 1935e, as
aeneipes
var.
sterbai
;
Radjabi 1968
;
Modarres Awal 1997
;
Borumand 2002
;
Holyński 2014
), southern provinces (
Abai & Adeli 1984
;
Modarres Awal 1997
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Afghanistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
),
Pakistan
(
Holyński 2014
).
*
Julodis amrishi
Makhan 2012: 1
.
Host plant(s) within
Iran
.
(Adult!)
Alhagi maurorum
(Fabaceae)
(Makhan 2012).
Distribution within
Iran
.
Markazi (Makhan 2012).
Distribution outside
Iran
.
None recorded.
Note.
Most probably a synonym of
J. partha
Obenberger 1923
(V. Kubáň, personal communication).
Julodis andreae
(
Olivier) 1790
: 53 (
Buprestis
);
Théry 1925
: 79
(variety of
onopordi
);
Salavatian 1950
: 7
(as
onopordi
var.
xanthographa
Faldermann
);
Radjabi 1968
: 73
(as
onopordi
andreae
and
onopordi sovitzi
), 1974: 39 (as
andreae
and
onopordi algirica
);
Bílý 1983a
: 31
, 32 (subspecies of
onopordi
and
onopordi sovitzi
);
Modarres Awal 1997
: 135
(as
andreae
), 136 (as
onopordi
,
onopordi
andreae
,
onopordi sovitzi
,
onopordi algirica
, and
sulcata
);
Borumand 2002
: 9
(as
onopordi algirica
), 10 (as subspecies of
onopordi
and
onopordi sovitzi
);
Kubáň & Volkovitsh 2006
: 326
;
Bellamy 2008a
: 49
; Sakenin
et al
. 2008: 31;
Havaskary
et al
. 2012
: 1
;
Ghobari
et al
. 2012
: 233
;
Radjabi & Gigli 2012
: 1
, 2 (as
andreae andreae
and
pietzchmanni
).
synonym:
sovitzii
(
Steven) 1830
: 158 (
Buprestis
);
Radjabi 1968
: 73;
Bílý 1983a
: 31 (subspecies of
onopordi
); Kubáň 2006: 47 (synonym of
andreae andreae
).
synonym:
setosa
(
Steven) 1830
: 156 (
Buprestis
, preoccupied by
setosa
Thunberg 1787
). synonym:
lineigera
Marseul 1865
: 62.
synonym:
ampliata
Marseul 1865
: 66.
synonym:
stevensii
Saunders 1871
: 5 (
nomen novum
for
setosa
Steven 1830
).
synonym:
hampei
Thomson 1878
: 11.
synonym:
lineata
Kerremans 1886
: xxxvii.
synonym:
subviolacea
Abeille de Perrin 1904: 208 (variety of
onopordi
); Kubáň 2006: 51 (available name;
lectotype
);
Kubáň & Volkovitsh 2006
: 326.
synonym:
tenuelineata
Abeille de Perrin 1904: 208 (variety of
onopordi
); Kubáň 2006: 51 (available name;
lectotype
);
Kubáň & Volkovitsh 2006
: 326.
synonym:
derasa
Abeille de Perrin 1904: 208 (variety of
onopordi
); Kubáň 2006: 51 (available name, synonym of
andreae andreae
);
Kubáň & Volkovitsh 2006
: 326.
synonym:
costifera
Obenberger 1919
: 142 (subspecies of
onopordi
); Kubáň 2006: 47 (synonym of
andreae andreae
).
misidentification:
pietzchmanni
:
Radjabi & Gigli 2012
: 2
(V. Kubáň, personal communication).
Host plant(s) within
Iran
.
(Adult!)
Acantholimon gulistanum
(Plumbaginaceae)
(Sakenin
et al
. 2008).
Distribution within
Iran
.
"
Armenia
Persica
" (
types
of
setosa
and
sovitzii
) (
Steven 1830
), "Chaldée persane, Plateau persan, Poucht-é-Kouh, El-Bourz, Louristan", multiple locations (
Théry 1925
). Alborz (
Radjabi 1974
), Ardabil, Hormozgan, Ilam, Markazi, Sistan & Baluchestan (
Radjabi 1974
;
Borumand 2002
as
onopordi algirica
and
scovitzi
), East Azarbaijan (
Radjabi 1968
;
Radjabi 1974
as
onopordi algirica
and
scovitzi
;
Modarres Awal 1997
as
onopordi
and
onopordi andrea
;
Borumand 2002
;
Havaskary
et al
. 2012
), Golestan (
Radjabi 1968
; Sakenin
et al
. 2008), Guilan, Kermanshah (
Radjabi 1974
;
Borumand 2002
as
onopordi algirica
and
scovitzi
; (
Modarres Awal 1997
as
onopordi
var.
algirica
and
onopordi andrea
), Hamadan (
Radjabi 1968
, as
onopordi sovitzi
varr.
xanthographa
,
carinulosa
;
Modarres Awal 1997
as
onopordi sovitzi
), Isfahan (
Bílý 1983a
as
onopordi sovitzi
;
Radjabi 1974
;
Borumand 2002
as
onopordi algirica
and
scovitzi
), Khuzestan (
Salavatian 1950
as
onopordi
var.
xanthographa
;
Radjabi 1974
;
Borumand 2002
;
Radjabi & Gigli 2012
, as
andreae andreae
and
pietzchmanni
), Kordestan, West Azarbaijan (
Radjabi 1974
;
Bílý 1983a
;
Borumand 2002
as
onopordi algirica
and
scovitzi
;
Ghobari
et al
. 2012
), Lorestan (
Salavatian 1950
as
onopordi
var.
xanthographa
;
Radjabi 1968
, as
onopordi sovitzi
varr.
xanthographa
,
carinulosa
;
Radjabi 1974
;
Modarres Awal 1997
as
onopordi
var.
algirica
and
onopordi sovitzi
;
Borumand 2002
as
onopordi algirica
and
scovitzi
), Qazvin (
Bílý 1983a
;
Borumand 2002
;
Ghobari
et al
. 2012
), Tehran (
Salavatian 1950
as
onopordi
var.
xanthographa
;
Radjabi 1968
, as
onopordi sovitzi
varr.
xanthographa
,
carinulosa
;
Radjabi 1974
;
Modarres Awal 1997
, as
andrea
,
onopordi
var.
algirica
,
onopordi andrea
, and
onopordi sovitzi
;
Borumand 2002
as
onopordi algirica
and
scovitzi
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Armenia
,
Azerbaijan
,
Iraq
,
Syria
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Julodis armeniaca
Marseul 1865
: 80
;
Théry 1925
: 79
;
Radjabi, 1974
: 39
(as
var.
cyrus
);
Modarres Awal 1997
: 136
(as
var.
cyrus
);
Borumand 2002
: 1
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 51
;
Ghobari
et al
. 2012
: 232
;
Ghahari & Bellamy 2012
: 43
.
synonym:
orientalis
Kerremans 1899
: 272
.
synonym:
cyrus
Obenberger 1923a
: 19;
Radjabi 1974
: 39;
Modarres Awal 1997
: 136; Kubáň 2006: 47 (synonym of
armeniaca
).
Distribution within
Iran
.
"Chaldée persane: Aiemabad" (
Théry 1925
). Bushehr, Golestan (
Borumand 2002
), East Azarbaijan (
Ghahari & Bellamy 2012
), Kermanshah (
Radjabi 1974
;
Borumand 2002
), Kordestan (
Ghobari
et al
. 2012
), Mazandaran (
Modarres Awal 1997
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Armenia
,
Cyprus
,
Iraq
,
Syria
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
Records for
Algeria
and
Egypt
in
Bellamy (2008a)
are erroneous.
Julodis audouinii
Laporte & Gory 1835: 16
;
Théry 1925
: 78
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 52
;
Ghobari
et al
. 2012
: 233
.
synonym:
theryi
Abeille de Perrin 1900: 3; Kubáň 2006: (synonym of
audouinii
). Biology/Bionomy:
Holdhaus 1920
: 50.
Distribution within
Iran
.
Kordestan (
Ghobari
et al
. 2012
), Lorestan (
Théry 1925
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Iraq
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Julodis balucha
Obenberger 1923a
: 17
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 52
;
Holyński 2014
: 18
. synonym:
semenovi
Alexeev
in
Alexeev
et al
. 1990
: 63
(subspecies of
intricata
); Kubáň 2006: 47 (synonym of
balucha
).
Distribution within
Iran
.
Khorasan (
Alexeev
et al
. 1990
, as
intricata
semenovi
;
Holyński 2014
), Sistan & Baluchestan (
Obenberger 1923a
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Afghanistan
,
Pakistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
;
Holyński 2014
).
*
Julodis cailliaudi
(
Latreille) 1827
: 277
(
Buprestis
);
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 54
(
caillaudi
[sic!]); Kubáň
in
Bílý
et al
. 2011
: 170
;
Ghahari & Bellamy 2012
: 43
(
caillaudi
).
synonym:
aethiopica
Laporte 1835: 162;
Kubáň & Volkovitsh 2006
: 327;
Bellamy, 2008a
: 54 (synonym of
caillaudi
[sic!]); Kubáň
in
Bílý
et al
. 2011
: 170.
Distribution within
Iran
.
Bushehr (
Ghahari & Bellamy 2012
),?southern
Iran
(
Jakobson 1913
; Kubáň
in
Bílý
et al
. 2011
).
Distribution outside
Iran
.
?
Algeria
,
Cameroon
,
Central African Republic
,
Chad
,
Democratic Republic of Congo
,
Egypt
,
Eritrea
,
Ethiopia
,
Gambia
,?
Libya
,
Mali
,
Mauritania
,
Niger
,
Nigeria
,
Senegal
,
Sudan
,?
Tunisia
(Kubáň
in
Bílý
et al
. 2011
).
Note.
The occurrence of this species in
Iran
needs confirmation. Previously it was indicated for
Iran
by
Jakobson (1913)
who cited the old publication of
Osculati (1844)
.
Julodis consobrina
Kerremans 1914
: 247
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 59
. synonym:
hauseri
Obenberger 1928b
: 65
;
Radjabi 1968
: 72
(subspecies of
gotwendensis
);
Modarres Awal 1997
: 136
(subspecies of
gotwendensis
); Kubáň 2006: 47 (synonym of
consobrina consobrina
).
Distribution within
Iran
.
Bushehr (
Obenberger 1928b, as
hauseri
),
Fars
(
Modarres Awal 1997
, as
gotwendensis
hauseri
),
Iran
(no locality cited) (
Kerremans 1914
;
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Julodis consobrina gotwendensis
Obenberger 1924c
: 6
(subspecies of
ethema
);
Bílý 1983a
: 31
;
Borumand 2002
: 2
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 59
; Sakenin
et al
. 2008: 32.
misidentification:
faldermanni eoa
:
Bílý 1983a
: 31
(not
Obenberger 1923a
) (V. Kubáň, personal communication).
Host plant(s) within
Iran
.
(Adult!)
Anchusa officinalis
(Boraginaceae)
(Sakenin
et al
. 2008).
Distribution within
Iran
.
Bushehr, Hormozgan (
Bílý 1983a
;
Borumand 2002
),
Fars
(
Radjabi 1968
: 72;
Borumand 2002
), Kerman, Kermanshah, Sistan & Baluchestan, Tehran (
Borumand 2002
), Khorasan (Sakenin
et al
. 2008), Khuzestan (
Obenberger 1924c
;
Borumand 2002
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
According to V. Kubáň (personal communication), part of specimens identified as
J. c. gotwendensis
(Loc. no. 308),
J. euphratica beludjistana
and
J. e. euphratica
(both from Loc. no. 304) in
Bily (1983a)
actually belong to
J
.
consobrina consobrina
. The records for Kerman, Kermanshah, Sistan & Baluchestan, Tehran (
Borumand 2002
), and Khorasan (Sakenin
et al
. 2008) are most probably misidentifications with different subspecies of
J. euphratica
. Additionaly,
J
.
pietzehmanni
var.
deserticoloides
Théry 1926
(see below) is a junior synonym of
J. c. gotwendensis
. According to V. Kubáň,
J. c. consobrina
is a habitant of clay and stony soils on the hills while
J. c. gotwendensis
is associated with coastal sandy soils.
Julodis cylindrica
Théry 1925
: 82
(as
cylindricus
);
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 59
.
Distribution within
Iran
.
?Ilam ("Chaldée persane: Aivan-i-Kerkha, Neima") (
Théry 1925
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
This species is known only from
type
locality (V. Kubáň, personal communication).
Julodis ehrenbergii
Laporte 1835: 162
;
Bílý 1983a
: 32
(
ehrenbergi
, subspecies of
onopordi
);
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 61
.
synonym:
bohemanni
Mannerheim 1837
: 19;
Kubáň & Volkovitsh 2006
: 327.
Distribution within
Iran
.
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Albania
,
Bulgaria
,
Croatia
,
Cyprus
,
Egypt
(Sinai),
Greece
,
Iraq
,
Macedonia
,
Syria
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Julodis escalerae
Abeille
de Perrin 1904: 206;
Théry 1925
: 78
;
Radjabi 1968
: 72
;
Bílý 1983a
: 31
;
Modarres Awal 1997
: 136
;
Borumand 2002
: 3
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 61
;
Radjabi & Gigli 2012
: 1
, 2. synonym:
bouvieri
Kerremans 1905
: 227
.
Distribution within
Iran
.
"Chaldée persane: Neima" (
Théry 1925
). Golestan, Kermanshah (
Radjabi 1968
;
Modarres Awal 1997
;
Borumand 2002
), Ilam (
Borumand 2002
),?Kerman (
Bílý 1983a
), Khuzestan (
Radjabi 1968
;
Modarres Awal 1997
;
Borumand 2002
;
Radjabi & Gigli 2012
), Lorestan (
Bílý 1983a
),
Iran
(no locality cited) (Abeille de Perrin 1904;
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
According to V. Kubáň (personal communication), specimens from Kerman in
Bílý (1983a, Loc. no. 347)
belong to
J
.
matthiesseni
Reitter 1905
(see below); specimens indicated for Khuzestan (
Bílý 1983a, Locs. no. 285 and 286
) actually originated from Lorestan.
Julodis euphratica
Laporte & Gory 1835: 18
;
Théry 1925
: 78
(as
iris
);
Salavatian 1950
: 6
(as
euphratica
and
iris
var.
euphratica
);
Davatchi
et al
. 1959
: 243
(variety of
iris
);
Farahbakhsh 1960
: 74
(variety of
iris
);
Adeli 1972
: 11
(variety of
iris
);
Radjabi 1968
: 72
, 1976: 128;
Bílý 1983a
: 32
;
Abai & Adeli 1984
: 57
(variety of
iris
), 116 (as
euphratica
);
Modarres Awal 1997
: 136
(as
euphratica
,
iris
and
iris
var.
euphratica
);
Faghih 1998
: 184
;
Borumand 2002
: 3
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 61
; Sakenin
et al
. 2008: 33;
Bílý
et al
. 2011
: 173
;
Havaskary
et al
. 2012
: 1
;
Ghobari
et al.
2012
: 233
;
Ghahari & Bellamy 2012
: 43
.
Host plant(s) within
Iran
.
Pistacia
roots (
Anacardiaceae
) (
Farahbakhsh 1960
;
Adeli 1972
), (Adult!)
Pistacia
spp., bark (
Abai & Adeli 1984
), pistachio, tamarisk (
Tamarix
sp.,
Tamaricaceae
) (
Modarres Awal 1997
),
Tamarix
spp., sapwood (
Abai & Adeli 1984
),
Cerasus vulgaris
(Rosaceae)
(Sakenin
et al
. 2008).
Distribution within
Iran
.
Bushehr, Qazvin (
Bílý 1983a
), Chaharmahal & Bakhtiari, Hormozgan, Lorestan, Mazandaran, Qom, Semnan, Sistan & Baluchestan, Yazd, Zanjan (
Borumand 2002
), East Azarbaijan (
Havaskary
et al
. 2012
),
Fars
(
Salavatian 1950
;
Davatchi
et al.
1959
;
Farahbakhsh 1960
;
Radjabi 1976
;
Modarres Awal 1997
;
Borumand 2002
;
Ghahari & Bellamy 2012
), Golestan (
Davatchi
et al
. 1959
;
Farahbakhsh 1960
), Isfahan (
Salavatian 1950
;
Davatchi
et al
. 1959
;
Farahbakhsh 1960
;
Adeli 1972
;
Bílý 1983a
;
Abai & Adeli 1984
;
Modarres Awal 1997
;
Faghih 1998
;
Borumand 2002
), Kerman (
Adeli 1972
;
Bílý 1983a
;
Borumand 2002
), Khorasan (
Salavatian 1950
;
Bílý 1983a
;
Modarres Awal 1997
;
Borumand 2002
; Sakenin
et al
. 2008), Kordestan (
Ghobari
et al
. 2012
), Tehran (
Modarres Awal 1997
, as
iris
;
Borumand 2002
), southern provinces (
Abai & Adeli 1984
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Afghanistan
,
Egypt
(Sinai),
Iraq
,
Jordan
,
Kyrgyzstan
,
Oman
,
Saudi Arabia
,
Syria
,
Tajikistan
,
Turkey
,
Turkmenistan
,
UAE
,
Uzbekistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
;
Bílý
et al
. 2011
).
Note.
According to V. Kubáň (personal communication),
J
.
euphratica euphratica
may occur only in the westernmost lowlands of
Iran
(Ilam, Khuzestan, northern Bushehr, western Lorestan); this subspecies is distributed in Arabian Peninsula and lowland
Iraq
(
Bílý
et al
. 2011
).
*
Julodis euphratica beludjistana
Obenberger 1924b
: 6
;
Radjabi 1968
: 72
;
Bílý 1983a
: 32
;
Modarres Awal 1997
: 136
;
Borumand 2002
: 5
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 62
;
Hadian
et al
. 2012
: 2
;
Holyński 2014
: 32
(subspecies of
whitehilli
Gray 1832).
Distribution within
Iran
.
Bushehr (
Radjabi 1968
;
Bílý 1983a
), Hormozgan (
Radjabi 1968
;
Bílý 1983a
;
Borumand 2002
), Isfahan, Khorasan, Khuzestan, Lorestan, Qazvin, Qom (
Borumand 2002
), Kerman, Sistan & Baluchestan (
Radjabi 1968
;
Bílý 1983a
;
Modarres Awal 1997
;
Borumand 2002
), Semnan (
Hadian
et al
. 2012
).
Distribution outside
Iran
.
Pakistan
(
Kubáň & Volkovitsh 2006
).
Note.
Probably misidentification, this subspecies known only from
Pakistan
. According to V. Kubáň (personal communication), all records in
Bílý (1983a)
refer to
J. e. scenica
Kerremans 1905
, record for Semnan (
Hadian
et al
. 2012
) and records for western, northern and central
Iran
most probably refer to
J. e. karelinii
Mannerheim 1837
.
Julodis euphratica bushirensis
Obenberger 1928b
: 66
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 62
.
Distribution within
Iran
.
Bushehr (
Obenberger 1928b
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Julodis euphratica karelinii
Mannerheim 1837
: 123
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 62
.
Distribution within
Iran
.
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Tajikistan
,
Turkmenistan
,
Uzbekistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
According to V. Kubáň (personal communication), the records for
J. euphratica beludjistana
Obenberger 1924
from western, northern and central
Iran
most probably refer to this subspecies.
*
Julodis euphratica nusskina
Obenberger 1924c
: 6
(subspecies of
euphratica
);
Radjabi 1968
: 72
;
Bílý 1983a
: 32
(synonym of
euphratica beludjistana
);
Modarres Awal 1997
: 136
(variety of
euphratica
);
Borumand 2002
: 5
(synonym of
euphratica beludjistana
);
Kubáň & Volkovitsh 2006
: 327
(subspecies of
euphratica
);
Bellamy 2008a
: 62
(subspecies of
euphratica
);
Holyński 2014
: 32
(synonym of
whitehilli
beludjistana
Obenberger 1924
).
Distribution within
Iran
.
See under
J. euphratica beludjistana
).
Distribution outside
Iran
.
Pakistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
J. nusskina
sensu
Bílý 1983a
(not
Obenberger 1924c
) (synonym of
euphratica beludjistana
sensu
Bílý 1983a
, not
Obenberger 1924c
) corresponds to
J. euphratica scenica
(V. Kubáň, personal communcation). This subspecies is known only from
Afghanistan
and
Pakistan
.
Julodis euphratica scenica
Kerremans 1905
: 231
;
Bílý 1983a
: 32
(as
klapaleki
and
euphratica beludjistana
);
Borumand, 2002
: 7
(as
klapaleki
);
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 62
.
synonym:
ormarensis
Obenberger 1924c
: 10;
Salavatian 1950
: 6;
Radjabi 1968
: 73;
Modarres Awal 1997
: 136;
Kubáň & Volkovitsh 2006
: 45 (new assignment), 327;
Holyński 2014
: 30 (subspecies of
whitehilli
Gray 1832).
Host plant(s) within
Iran
.
(Adult!)
Ononis
(Fabaceae)
(
Salavatian 1950
;
Modarres Awal 1997
, as
ormarensis
).
Distribution within
Iran
.
Kerman (as
ormarensis
) (
Salavatian 1950
;
Modarres Awal 1997
), Sistan & Baluchestan (as
klapaleki
) (
Bílý 1983a
;
Borumand 2002
), (as
ormarensis
) (
Salavatian 1950
;
Radjabi 1968
;
Modarres Awal 1997
),
Iran
(no locality cited) (
Kerremans 1905
;
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
: 62).
Distribution outside
Iran
.
Pakistan
(
Kubáň & Volkovitsh 2006
;
Bellamy, 2008a
).
Note.
According to V. Kubáň (personal communication), the records for
J. euphratica beludjistana
Obenberger 1924
from South-East
Iran
refer to this subspecies.
Julodis faldermanni
Mannerheim 1837
: 14
;
Bílý 1983a
: 31
(as
variolaris variolaris
);
Modarres Awal 1997
: 136
;
Sakenin
et al
. 2000
: 301
;
Kubáň & Volkovitsh 2006
: 327
;
Bellamy 2008a
: 62
.
misidentification:
variolaris variolaris
:
Bílý 1983a
: 31
(part.) (V. Kubáň, personal communication).
Host plant(s) within
Iran
.
(Adult!)
Zygophyllum
sp. (
Zygophyllaceae
) (
Sakenin
et al
. 2000
).
Distribution within
Iran
.
Fars
(
Modarres Awal 1997
), Semnan (
Sakenin
et al
. 2000
), West Azarbaijan (
Bílý 1983a, as
variolaris variolaris
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
).
Distribution outside
Iran
.
Armenia
,
Azerbaijan
,
Georgia
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
The records for
Fars
and Semnan in
Modarres Awal (1997)
and
Sakenin
et al
. (2000)
, and
Iraq
in
Bellamy (2008a)
are most probably based on misidentifications.
Julodis intricata
Redtenbacher 1843
: 982
;
Théry 1925
: 78
;
Radjabi 1974
: 39
;
Modarres Awal 1997
: 136
;
Borumand 2002
: 7
;
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 69
;
Ghobari
et al
. 2012
: 233
. synonym:
ruginota
Marseul 1865
: 54
.
synonym:
cupreocoelata
Marseul 1865
: 53.
synonym:
rugosa
Thomson 1878
: 9 (preoccupied by
rugosa
Laporte & Gory 1835); Kubáň 2006: 45 (new assignment).
synonym:
thomsoni
Kerremans 1892
: 23 (
nomen novum
for
rugosa
Thomson 1878
).
Distribution within
Iran
.
Kermanshah (
Radjabi 1974
;
Modarres Awal 1997
;
Borumand 2002
), Kordestan (
Ghobari
et al
. 2012
), Lorestan ("Tchéar-Dooul") (
Théry 1925
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Iraq
,
Israel
,
Syria
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Julodis laevicostata
Gory 1840
: 16
;
Théry 1925
: 79
;
Salavatian 1950
: 7
;
Radjabi 1968
: 72
, 1976: 128;
Bílý 1983a
: 32
;
Modarres Awal 1997
: 136
;
Borumand 2002
: 8
;
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 70
;
Ghobari
et al
. 2012
: 233
.
Distribution within
Iran
.
"Azerbedjan: Massif du Sahend; Elbourz: Haute Valée de Charoud" (
Théry 1925
). Bushehr, Chaharmahal & Bakhtiari, East Azarbaijan, Isfahan, Kerman, Kordestan, Semnan, West Azarbaijan (
Borumand 2002
),
Fars
(
Radjabi 1976
;
Borumand 2002
), Khorasan, Lorestan (
Salavatian 1950
;
Borumand 2002
), Kuhgiloyeh & Boyerahmad, Mazandaran (
Bílý 1983a
), Tehran (
Salavatian 1950
;
Bílý 1983a
;
Borumand 2002
), generally distributed (
Radjabi 1968
,
Modarres Awal 1997
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Iraq
,
Turkmenistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
*
Julodis longicollis
Abeille de Perrin 1904: 207 (variety of
onopordi
).
Kubáň & Volkovitsh 2006
: 329
(species incertae sedis; distribution unknown);
Bellamy 2008a
: 71
.
Distribution within
Iran
.
Iran
(no locality cited) (
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
The occurrence of this poorly known species in
Iran
needs confirmation.
Julodis matthiesseni
Reitter 1905
: 204
;
Théry 1925
: 78
;
Radjabi 1974
: 39
;
Borumand 2002
: 9
;
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 72
.
misidentification:
escalerae
:
Bílý 1983a
: 31
(not Abeille de Perrin 1904, part.) (see above) (V. Kubáň, personal communication).
Distribution within
Iran
.
"Chaldée persane: Kerkha au Kébir Kouh" (
Théry 1925
). Bushehr, Kermanshah (
Radjabi 1974
;
Borumand 2002
), Kerman (
Radjabi 1974
;
Bílý 1983a
, as
escalerae
;
Borumand 2002
), central
Iran
(
Reitter 1905
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
According to V. Kubáň (personal communication), records for Bushehr and Kermanshah in
Radjabi (1974)
and
Borumand (2002)
are most probably misidentifications of
J. punctatocostata
Gory 1840
(Bushehr)
and
J. laevicostata
Gory 1840
(Kermanshah)
.
Julodis partha
Obenberger 1923a
: 23
(subspecies of
onopordi
);
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 75
.
synonym:
straussi
Pochon 1967
: 205
;
Radjabi 1974
: 39;
Modarres Awal 1997
: 136;
Borumand 2002
: 12; Kubáň 2006: 47 (synonym of
partha
).
Distribution within
Iran
.
East Azarbaijan, Kordestan (
Radjabi 1974
;
Modarres Awal 1997
;
Borumand 2002
, as
J. straussi
Pochon 1967
), Lorestan (
Pochon 1967, as
straussi
),
Iran
(no locality cited) (
Obenberger 1923a
;
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Julodis pietzchmanni
Kerremans 1914
: 249
;
Radjabi 1974
: 39
;
Modarres Awal 1997
: 136
;
Faghih 1998
: 184
;
Borumand 2002
: 3
;
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 76
;
Radjabi & Gigli 2012
: 1
, 3 (as
aeneipes
). synonym:
deserticoloides
Théry 1926: 39
(variety of
pietzehmanni
);
Radjabi 1974
: 39
, 1976: 128;
Modarres Awal 1997
: 136
;
Borumand 2002
: 3
.
misidentification:
aeneipes
:
Radjabi & Gigli 2012
: 3
(V. Kubáň, personal communication).
Distribution within
Iran
.
Bushehr, Kuhgiloyeh & Boyerahmad (
Radjabi 1974
;
Borumand 2002
, as
J. deserticoloides
),
Fars
(
Radjabi 1974
,
1976
;
Modarres Awal 1997
,
Borumand 2002
, as
J. deserticoloides
), Isfahan (
Faghih 1998 as
J. deserticoloides
), Khuzestan (
Radjabi 1974
;
Modarres Awal 1997
;
Borumand 2002
, as
J. deserticoloides
;
Radjabi & Gigli 2012
: as
aeneipes
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Iraq
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
According to V. Kubáň (personal communication)
J. pietzehmanni
var.
deserticoloides
is a junior synonym of
J. consobrina gotwendensis
. It is possible that all records of
J. pietzchmanni
from
Iran
refer to the latter (Bushehr, Khuzestan, Kuhgiloyeh & Boyerahmad, Isfahan) and
J. aeneipes
(
Fars
)
.
Julodis proxima
Gory 1840
: 12
;
Théry 1925
: 83
(variety of
iris
);
Radjabi 1968
: 73
, 1976: 128 (subspecies of
euphratica
);
Modarres Awal 1997
: 136
(as
euphratica
proxima
and
proxima
);
Borumand 2002
: 6
(subspecies of
euphratica
); Kubáň 2006: 43 (full species);
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 77
;
Radjabi & Gigli 2012
: 1
, 3 (as
euphratica
);
Holyński 2014
: 23
(subspecies of
whitehilli
Gray 1832).
synonym:
beesoni
Obenberger 1934e
: 186; Kubáň 2006: 47 (synonym of
proxima
).
misidentification:
euphratica
:
Radjabi & Gigli 2012
: 3
(V. Kubáň, personal communication).
Distribution within
Iran
.
Fars
(
Radjabi 1968
,
1976
;
Modarres Awal 1997
, as
euphratica
proxima
), Hormozgan, Ilam, Sistan & Baluchestan, Tehran (
Borumand 2002
),?Ilam ("Chaldée persane: Dehleroum") (
Théry 1925; loc. no. 8
), Khuzestan (
Radjabi & Gigli 2012, as
euphratica
), generally distributed except Caspian Sea area (
Modarres Awal 1997
, as
proxima
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Iraq
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
According to V. Kubáň (personal communication) most records for
J. iris
(except for loc. no. 8) in
Théry (1925)
refer to
J. euphratica bushirensis
Obenberger 1928
and
J. euphratica karelinii
Mannerheim 1837
; the records for
Fars
, "generally distributed except Caspian Sea area" in
Radjabi (1968)
and
Modarres Awal (1997)
, and Hormozgan, Ilam, Sistan & Baluchestan, Tehran in
Borumand (2002)
are most probably misidentifications, this rare species occurs only in Western
Iran
.
Julodis punctatocostata
Gory 1840
: 17
; Kubáň 2006: 43 (full species);
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 78
.
synonym:
quadricostata
Marseul 1865
: 71; Kubáň 2006: 47 (synonym of
punctatocostata
).
Distribution within
Iran
.
Iran
(no locality cited) (
Gory 1840
;
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Julodis punctatocostata ramifera
Marseul 1865
: 69
;
Radjabi 1968
: 73
(subspecies of
laevicostata
);
Modarres Awal 1997
: 136
(subspecies of
laevicostata
); Kubáň 2006: 43 (subspecies of
punctatocostata
);
Kubáň & Volkovitsh 2006
: 328
;
Bellamy 2008a
: 78
.
Distribution within
Iran
.
East Azarbaijan (
Radjabi 1968
;
Modarres Awal 1997
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
According to V. Kubáň (personal communication)
J. p.
punctatocostata
(Western
Fars
) and
J. p. ramifera
("Persepolis") are known from
Fars
; the record for East Azarbaijan in
Radjabi (1968)
and
Modarres Awal (1997)
refer to
J. laevicostata
Gory 1840
.
Julodis speculifer
Laporte 1835: 160
;
Bílý 1983a
: 31
(as
speculifera
),
Radjabi 1968
: 73
(as
speculifera
);
Modarres Awal 1997
: 136
(as
speculifera
);
Borumand 2002
: 11
(as
speculifera
);
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 79
.
Distribution within
Iran
.
Bushehr (
Radjabi 1968
;
Bílý 1983a
;
Borumand 2002
),
Fars
(
Radjabi 1968
;
Modarres Awal 1997
), Hormozgan, Khuzestan (
Radjabi 1968
;
Modarres Awal 1997
;
Borumand 2002
), Ilam, Kermanshah (
Borumand 2002
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Iraq
(
s.
speculifer
),
Jordan
,
Kuwait
,
Saudi Arabia
(
s. disconae
Théry, 1936a
) (
Kubáň & Volkovitsh 2006
).
Julodis speculifer monstrosa
Abeille
de Perrin 1904: 206;
Radjabi 1968
: 73
;
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 80
;
Radjabi & Gigli 2012
: 1
, 2.
synonym:
iranica
Obenberger 1923a
: 17
.
Distribution within
Iran
.
"Gotvend" (Abeille de Perrin 1904). Bushehr (
Obenberger 1923a, as
iranica
), Ilam, Kermanshah (
Radjabi 1968
), Khuzestan (
Radjabi & Gigli 2012
);
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.
Note.
The records for
Egypt
,
Israel
,
Syria
, and
Turkey
in
Bellamy (2008a)
are dubious.
*
Julodis syriaca
(
Olivier) 1790
: no 32: 87 (
Buprestis
);
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 81
. synonym:
philistina
Obenberger 1934e
: 187
.
Distribution outside
Iran
.
Egypt
,
Iraq
,
Israel
,
Jordan
,
Lebanon
,
Syria
,
Turkey
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
*
Julodis syriaca distincta
Gory 1840
: 11
;
Théry 1925
: 78
(variety of
speculifera
);
Salavatian 1950
: 70
(species);
Modarres Awal 1997
: 136
(species); Kubáň 2006: 43 (subspecies of
syriaca
);
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 81
.
synonym:
loeffleri
Mandl & Pochon 1957: 323.
Distribution within
Iran
.
"Chaldée persane: locs. no. 4, 5, 8, 9; Poucht-é-Kouh: loc. no. 24" (
Théry 1925
). Kermanshah, Khuzestan (
Salavatian 1950
;
Modarres Awal 1997
).
Distribution outside
Iran
.
Iraq
,
Syria
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
The records for
Iran
need confirmation.
Julodis variolaris
(
Pallas) 1771
: 464 (
Buprestis
);
Farahbakhsh 1960
: 75
;
Abai & Adeli 1984
: 68
;
Modarres Awal 1997
: 137
;
Borumand 2002
: 12
;
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 82
; Makhan
et al
. 2012: 1;
Holyński 2014
: 34
.
synonym:
kirghisica
Marseul 1865
: 47 (variety of
variolaris
).
synonym:
oberthuri
Kerremans 1898a
: 134;
Kubáň & Volkovitsh 2006
: 44 (new assignment);
Holyński 2014
: 23 (subspecies of
whitehilli
Gray 1832).
synonym:
seminata
Abeille de Perrin 1904: 211.
synonym:
amurensis
Abeille de Perrin 1904: 211.
Host plant(s) within
Iran
.
Larvae attack roots Pistachio (
Pistacia
,
Anacardiaceae
), lucerne (
Medicago
),
Alhagi
(Fabaceae)
(
Farahbakhsh 1960
;
Modarres Awal 1997
),
Pistacia
spp., bark (
Abai & Adeli 1984
),
Alhagi maurorum
(Fabaceae)
(Makhan
et al
. 2012).
Distribution within
Iran
.
Fars
, Isfahan (
Farahbakhsh 1960
;
Abai & Adeli 1984
;
Modarres Awal 1997
), Khorasan (
Bílý 1983a
;
Borumand 2002
, as subspecies
freygessneri
Meyer-Darcis 1883
), Kerman,?West Azarbaijan (
Bílý 1983a
), Semnan (Makhan
et al
. 2012),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Afghanistan
,
Azerbaijan
,
China
,
Kazakhstan
,
Kyrgyzstan
,
Russia
,
Tajikistan
,
Turkmenistan
,
Uzbekistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
Specimens of
J. variolaris variolaris
from West Azarbaijan (Loc. no. 267) in
Bílý (1983a)
belong to
J. faldermanni
(see above) (V. Kubáň, personal communication).
Julodis variolaris brendelli
Holyński 1984
: 524
(subspecies of
variolaris
);
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 83
.
Distribution within
Iran
.
Fars
(
Holyński 1984
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
: 83).
Distribution outside
Iran
.
None recorded.
*
Julodis variolaris eoa
Obenberger 1923a
: 15
;
Salavatian 1950
: 7
;
Radjabi 1968
: 72
;
Bílý 1983a
: 31
;
Modarres Awal 1997
: 136
(as species);
Borumand 2002
: 7
(subspecies of
faldermanni
);
Kubáň & Volkovitsh 2006
: 329
(subspecies of
variolaris
);
Bellamy 2008a
: 83
;
Holyński 2014
: 34
.
Host plant(s) within
Iran
.
(Adult!) Milk vetch (
Astragalus
sp.,
Fabaceae
) (
Salavatian 1950
;
Modarres Awal 1997
).
Distribution within
Iran
.
Bushehr (
Bílý 1983a
), Golestan, Kerman, Sistan & Baluchestan (
Radjabi 1968
;
Bílý 1983a
;
Modarres Awal 1997
;
Borumand 2002
), Hormozgan, Khorasan, Mazandaran, Semnan (
Borumand 2002
), Tehran (
Salavatian 1950
;
Radjabi 1968
;
Modarres Awal 1997
;
Borumand 2002
), Kermanshah (
Radjabi 1968
;
Modarres Awal 1997
), southeastern
Iran
(
Holyński 2014
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
Afghanistan
,
Pakistan
(
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Note.
According to V. Kubáň (personal communication)
J. variolaris eoa
in
Bílý (1983a, Loc. no. 308)
actually belongs to
J. consobrina gotwendensis
. Occurrence of this subspecies in
Iran
needs confirmation.
*
Julodis variolaris freygessneri
Meyer-Darcis 1883
: xxxix;
Farahbakhsh 1960
: 75
;
Abai & Adeli 1984
: 68
;
Modarres Awal 1997
: 137
;
Borumand 2002
: 12
;
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 83
. Immatures:
Bílý 1983b
: 65
.
synonym:
undulata
Heyden 1883: 107
.
Host plant(s) within
Iran
.
Pistachio roots (
Pistacia
sp.,
Anacardiaceae
) (
Farahbakhsh 1960
;
Modarres Awal 1997
), bark (Adult!) (
Abai & Adeli 1984
).
Distribution within
Iran
.
Khorasan (
Borumand 2002
), Markazi, central provinces (
Abai & Adeli 1984
), northern provinces (
Farahbakhsh 1960
;
Modarres Awal 1997
),
Iran
(no locality cited) (
Bellamy 2008a
).
Distribution outside
Iran
.
Afghanistan
,
Turkmenistan
,
Uzbekistan
(
Bellamy 2008a
).
Note.
According to V. Kubáň (personal communication) all records for different subspecies of
J. variolaris
with the exception
J. v. brendelli
Holyński
from
Fars
and
J. v. eoa
Obenberger
from Sistan & Baluchestan refer to
J. v. variolaris
.
Julodis zablodskii
Motschulsky 1844
: 33
;
Kubáň & Volkovitsh 2006
: 329
;
Bellamy 2008a
: 85
.
Distribution within
Iran
.
Golestan (
Motschulsky 1844
),
Iran
(no locality cited) (
Kubáň & Volkovitsh 2006
;
Bellamy 2008a
).
Distribution outside
Iran
.
None recorded.