Cutting the Gordian Knot: Phylogenetic and ecological diversification of the Mesalina brevirostris species complex (Squamata, Lacertidae)
Author
Jiří Šmíd
Author
Jiří Moravec
Author
Václav Gvoždík
Author
Jan Štundl
Author
Daniel Frynta
Author
Petros Lymberakis
Author
Paschalia Kapli
Author
Thomas Wilms
Author
Andreas Schmitz
Author
Mohammed Shobrak
Author
Saeed Hosseinian Yousefkhani
Author
Eskandar Rastegar-Pouyani
Author
Aurora M. Castilla
Author
Johannes Els
Author
Werner Mayer
text
Zoologica Scripta
2017
2017-10-31
46
6
1
30
journal article
10.1111/zsc.12254
44b12099-59c0-4c00-b34c-c4f14d37ec56
1037920
Mesalina bernoullii
(Schenkel, 1901)
comb. nov.
Eremias bernoullii
– Schenkel (1901)
Eremias brevirostris
– Boulenger (1887)
Eremias brevirostris
forma
typica
– Angel (1936)
Eremias brevirostris fieldi
– Haas & Werner (1969).
Holotype
MCZ
56617
.
Type
locality
Mahor Birinji
,
Khuzestan Province
,
Iran
. MorphoBank pictures: M407313–M407317.
Mesalina brevirostris
– Szczerbak (1989)
Mesalina brevirostris brevirostris
– Szczerbak (1989)
Holotype
.
NMB 4396
.
Type
locality: ‘
Palmyra’
(
Syria
).
MorphoBank
pictures: M407230– M407235.
Distribution. Sinai,
Jordan
,
Syria
,
Iraq
, southwestern
Iran
,
Kuwait
, northeastern
Saudi Arabia
(
Fig. 1
).
Notes.
Mesalina bernoullii
represents a morphologically very variable species (
Table S5
). Several geographically isolated morphotypes that differ considerably in body size and scalation can be found within its large range. For example, Haas and Werner (1969) assigned a subspecific status (
Eremias brevirostris fieldi
) to the population from southwestern
Iran
characterized by low numbers of dorsal and gular scales and subdigital lamellae and Moravec (2004) described a morphotype from Jabal al Arab, southern
Syria
, that has unusually high numbers of scales of the same characters. Nevertheless, our results did not show any correlation between this morphological and phylogenetic clustering.
The material analyzed in this study included specimens from the
type
locality of
Eremias bernoullii
Schenkel, 1901
corresponding morphologically to the
holotype
of this taxon. It also included a specimen from ca.
40 km
of the
type
locality of
E. brevirostris fieldi
Haas and Werner, 1969
corresponding morphologically to this form, as well as individuals representing the Jabal al Arab morphotype. Based on the results of the phylogenetic and morphological analyses we conclude that the name
Eremias brevirostris fieldi
Haas and Werner, 1969
is a junior subjective synonym of
Eremias bernoullii
Schenkel, 1901
.