The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand Author Seldon, David S. Author Buckley, Thomas R. text Zootaxa 2019 2019-05-06 4598 1 1 148 journal article 26919 10.11646/zootaxa.4598.1.1 475fd53e-c5c6-4a99-bb2e-cc2382f0171f 1175-5326 2668063 6BA5F086-1014-46B5-B6D5-77FE0CD07AEC Mecodema mohi sp. n. Figure 38 . Diagnosis: Mecodema mohi is different from other North Island Mecodema species by having: 1 , vertexal groove very shallow and broad, defined laterally by sparse punctures, medially usually without microsculpture; 2 , pronotum disc with distinct, small depressions either side of midline medially, some may have punctures close to pronotal foveae; 3 , distinctive shape of the apical portion of the penis lobe ( Fig. 38 PL). Description: Length 28–34.8 mm , pronotal width 7.5–9.93 mm , elytral width 8.4–11.5 mm . Colour of entire body matte to glossy black, coxae and femur may be bright red to dark reddish-brown. Head: Very broad and flat. Vertex ( Fig. 9 ) with fine lines forming an isodiametric pattern laterally; vertexal groove very shallow and broad, defined laterally by sparse punctures ( Fig. 38 ), medially usually without microsculpture; large supraorbital puncture bearing 6–7 setae; 2 poorly defined supraorbital grooves; frons slightly convex medially; frontoclypeal suture indistinct, tentorial pits ( Fig. 9 ) small; anterior area of clypeus with broad and shallow grooves (may be indistinct), 1 large setose puncture each side bearing 2–3 setae ( Fig. 38 ), central puncture bearing 2 setae (may be absent). Labrum rectangular, anterior edge straight with 2 proximate central setae, 2 setae each side evenly spaced (may have an extra pair). Mentum lobes rounded ( Fig. 12F ), median process very short and broad, strongly angled upward (60°), distinctly indentate ( Fig. 12A ); mentum setae absent. Submentum sclerite constriction broad with 6–8 widely spaced setae. Stipes with 2 basal setae. Gula pits small, suture defined, gula flat with lines variably present or absent. Gena ( Fig. 8 ) with rugose wrinkles forming an isodiametric pattern across entire area. Prothorax: Prothoracic carina broad the entire length, slightly crenulated with 8–15 setae each side ( Fig. 38 ), extended beyond anterior angle; posterior lateral sinuation ( Fig. 11 ) evidently carinate, slightly angled inward; pronotum broad and flat, overall shape broadly cordate; midline well-defined, anterior medial triangular impression and posterior medial diamond-shaped impression present, disc with fine transverse wrinkles extended laterally ( Fig. 38 ), distinct small depressions either side of midline medially, some specimens may have punctures proximate foveae; pronotal foveae broad and deep; anterior and posterior edges relatively straight. Prosternum flat with very fine transverse lines, anterior edge ridged; proepisternum microsculpture absent. Procoxal setae absent; protibia distally expanded and shovel-like ( Fig. 10B ). Elytra: Broad and moderately deflected laterally; humeral angle evenly convex ( Fig. 6B ); basal margin moderately curved and gently sloped to base, interval 1 extended to scutellum; lateral carina ( Fig. 11 ) narrow the entire length, extended to humeral angle; humerus with 4–5 setose punctures, varying distribution along angle (asymmetrical) ( Fig. 38 ); suture impressed; striae 1–4 impressed with very small obsolescent punctures, striae 5–8 with punctures visibly present, all punctures small and regularly distributed; intervals 1–4 flat, intervals 5–8 moderately convex, interval 9 strongly convex; interval microsculpture present as transverse lines; 7 th strial setal pattern with 3–4 setose punctures in anterior ½, 4–5 setose punctures in posterior ½, setose punctures large. Ventral surface: Mesepisternum sparsely punctured; metepisternum without microsculpture; setose punctures present on mesocoxae (2) and metacoxae (2). All abdominal ventrites finely lineate; ventrites 3–5 ( Fig. 2 ) with 1 setose puncture each side of midline; ventrite 6 with setae present: ♂ with 2–3 large setose punctures each side, apical edge straight between distantly spaced setae, ♀ with 3 large setose punctures each side, apical edge straight between distantly spaced setae; ventrites without lateral foveae. Anterior metaventrite process ( Fig. 2 ) an equalsided triangle with narrow and well-defined carina the entire length. Male genitalia: Apical portion of penis lobe asymmetrically hooked with a slight deflection to right of vertical axis (VV) ( Fig. 15C ), apical portion narrow (60° angle in vertical plane), ventral process as pronounced as dorsal process ( Fig. 38 PL), ventral process a narrow and sharply rounded point, pushed forward of vertical axis ( Fig. 3 ), apex curved upward (flattened and symmetrical) to form slightly broadened dorsal process ( Fig. 38 PL), this is flattened (not recurved), brief curve returned to shaft; ventral edge of penis lobe curve to right distinct in apical ½ (VV), shaft narrowed slightly before apex but broadened to base, overall length of penis lobe distinctly curved ventrally (LV) ( Fig. 16C ). Structures of the endophallus ( Fig. 4 ): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule greatly expanded with projection; setal (scales) coverage of the apical plate extensive (76–100%); size of left setose flange small; size of right setose flange very large. Left paramere basal lobe broadly triangular due to enlarged dorsal hump, moderately (45°) sloped to short and narrow arm; terminal lobe slightly broadened with tuft of long setae ( Fig. 38 LP); setae extended along apical portion to arm only; ventral edge straight. Right paramere narrowly triangular ( Fig. 38 RP), gradually narrowed to apex with a double row of long setae extended along apical ¾ of ventral edge. Female genitalia: Basal gonocoxite 1 short and broad, apically flattened, ventral surface wrinkled with a few rugose grooves, internal dorso-lateral carina with 2 setae proximate base of gonocoxite 2, 1 seta double-spaced posterad ( 3 in total). Gonocoxite 2 ( Fig. 5 ) bluntly rounded and squatly triangular, apically reflexed. Ramus long and narrow. Comments: Mecodema mohi is one of many similar large oconnori -like species inhabitating many North Island forest fragments. It is relatively abundant and can be easily found by rolling logs in Mohi Bush Scenic Reserve. Distribution: New Zealand , North Island, Hawke’s Bay, Maraetotara Plateau, Mohi Bush S.R. Holotype : NZAC male labelled. NEW ZEALAND , HB, Maraetotara Plateau , Mohi Bush S.R. By hand. 3 Jan 2015 . D.S. Seldon , J.L. Druitt , K.G. Seldon / HOLOTYPE Mecodema mohi n. sp. design. DS Seldon, TR Buckley 2018 [red label]. FIGURE 38. Mecodema mohi plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe. Paratypes : 1♂ , 2♀ , New Zealand H.B. Mohi Bush Waimarama 10.XI.1985 T.H. and J.M. Davies / V / T.H. Davies Collection / AI.10473 ( NZAC ) ; 4, NEW ZEALAND HB, Maraetotara Plateau , Mohi Bush S.R. , 10 Dec 2012 – 7 Jan 2013 , D.S. Seldon & D. Brown / pitfall traps , regen. podocarp-tawa canopy S39°51’24.5 , E176°53’44.7 ( AMNZ ) ; 1 [card mounted], Maraetotara Gorge , Hawkes Bay , 7.10.92 , J.I. Townsend [hw] ( NZAC ) . Etymology: Mecodema mohi is named for the type locality (Mohi Bush S.R.).