Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae) Author Selnekovič, Dávid D930D757-D929-4248-A29C-A857E0505345 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. david.selnekovic@uniba.sk Author Jäch, Manfred A. D749707A-8823-4110-8D51-BF5AF4E2820F Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria. manfred.jaech@nhm-wien.ac.at Author Kodada, Ján 6E88BFBB-8769-44EC-8285-29E357CEC064 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. jan.kodada@uniba.sk text European Journal of Taxonomy 2024 2024-09-11 957 1 229 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247 journal article 10.5852/ejt.2024.957.2651 2118-9773 13773385 9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3 Elmomorphus mazzoldii sp. nov. urn:lsid:zoobank.org:act: 50262137-2A82-4389-BBF3-6578FD0B0DA2 Figs 35–36 , 37B , 110C Differential diagnosis Elmomorphus mazzoldii sp. nov. ( Fig. 35 ) is characterised by having a plastron on the cranial surface (missing only on the semicircular glabrous area of the vertex), in anterior angles of the pronotum and along lateral sides of the elytra. Such a combination of characters is shared with E. ellipticus sp. nov. , E. corpulentus sp. nov. , E. montanus , E. paramontanus sp. nov. and E. prosternalis . Elmomorphus mazzoldii differs from the mentioned species (except E. ellipticus ) in the shape of the body, which is widest closely before elytral midlength in E. mazzoldii ( Fig. 35 ), but widest behind elytral midlength in the other species. Moreover, the body dimensions are smaller in E. mazzoldii , with TL up to 2.8 mm versus at least 2.8 in other mentioned species, except E. ellipticus . Elmomorphus mazzoldii most closely resembles E. ellipticus , from which it differs in the more strongly convex pronotum and elytra ( Fig. 35 ), narrower plastron bands on elytra, each covering less than ⅓ of elytral width (in elytral midlength), and the shape of the aedeagus with shorter and broader phallobase and shorter parameres, PhL/PrL: 1.67 (n =1) in E. mazzoldii versus PhL/PrL: 1.53 (n =1) in E. ellipticus ( Fig. 36 ). Bursa copulatrix with numerous scattered spines ( Fig. 37B ). Etymology The epithet is a proper noun in the genitive case honouring Paolo Mazzoldi, an Italian entomologist who kindly provided interesting material. Type material Holotype THAILAND – Loei Province • ♂ ; “ THAILAND : Loei Prov. Phu Kradung NP 1150 m , 29.12.1999 leg. P. Mazzoldi (17)”; NMW . Paratypes THAILAND – Loei Province • 1 ♀ ; same collection data as for holotype; NMW • 1 ♀ ; “ THAILAND : Loei Prov. Phu Kradung NP 1270 m 27.12.1999 leg. P. Mazzoldi (12)”; NMW . Type locality Thailand , Loei Province , Phu Kradueng National Park. Description Dimensions (mm): TL: ♂ 2.63 (n =1), ♀♀ 2.60–2.83 (n=2); PL: ♂ 0.67 (n= 1), ♀♀ 0.61–0.71 (n =2); PW: ♂ 1.23 (n= 1), ♀♀ 1.16–1.26 (n =2); EL: ♂ 2.03 (n= 1), ♀♀ 1.96–2.21 (n =2); EW: ♂ 1.40 (n=1), ♀♀ 1.33–1.46 (n= 2); PhL: 0.48 (n =1); PrL: 0.29 (n=1). Fig. 35. Elmomorphus mazzoldii sp. nov. , holotype, male (NMW), TL: 2.63 mm. Fig. 36. Elmomorphus mazzoldii sp. nov. , holotype (NMW), aedeagus. A . Lateral aspect. B . Ventral aspect. Scale bar: 0.1 mm. Body oval, strongly convex, widest around middle of elytra ( Fig. 35 ). Integument black; mouthparts, antennae, and legs reddish brown. Pubescence consisting of short, sparse, decumbent yellowish setae. Dorsal plastron present on head (except for semicircular area on vertex), anterior angles of pronotum, and lateral portions of elytra. Ventral plastron absent on prosternal process, median part of metaventrite, median part of abdominal ventrite 1 and antero-median part of ventrite 2. Fig. 37. Bursa copulatrix and vagina. A . Elmomorphus ellipticus sp. nov. , paratype from type locality (NMW). B . Elmomorphus mazzoldii sp. nov. , paratype from type locality (NMW). Scale bar: 0.1 mm. Dorsal surface of head with round setiferous punctures, each slightly smaller than eye facet, separated by ca 1.0–1.5 × puncture diameters. Plastron present on dorsal surface, except on semicircular area on vertex reaching to midlength of eyes. Labrum transverse, anterior margin straight, exposed portion microreticulate with setiferous punctures. Anterior margin of clypeus straight. Eyes oval, protruding, ID: ♂ 0, 46 mm (n= 1), ♀♀ 0.41–0.47 mm (n =2). Antennae 9-segmented, densely setose. Pronotum transverse, moderately convex, PW/PL ratio: ♂ 1.82 (n= 1), ♀♀ 1.76–1.90 (n =2); lateral pronotal sides convergent, rounded; anterior angles strongly deflexed, protruding; surface smooth with round setiferous punctures ca as wide as those on elytra; plastron on anterior pronotal angles, posteriorly reaching pronotal midlength. Prosternal process without clusters of long setae, with lateral margins straight, posterior margin rounded, lateral portions weakly raised, median keel nearly flat. Scutellum longer than wide, smooth, with several setiferous punctures. Median part of metaventrite slightly convex, without clusters of long setae. Elytra oval, strongly convex, widest closely before midlength, EL/EW: ♂ 1.44 (n=1), ♀♀ 1.48–1.52 (n= 2); surface smooth with scattered, round setiferous punctures; plastron confined to lateral portions, anteriorly not reaching humeral callus, weakly expanding posteriad, covering less than ⅓ of elytron width (at elytral midlength). Tibiae straight; protibia as long as protarsus; PrTiL/PL: ♂ 1.04 (n=1), ♀♀ 1.00–1.03 (n =2). Terminal protarsomere as long as all preceding segments combined. Abdominal ventrites covered with plastron except for middle of ventrite 1 and antero-median portion of ventrite 2. Ventrite 1 with two indistinct admedian keels. Male ventrite 5 narrowly truncate at apex; female ventrite 5 with short apical keel. Aedeagus ( Fig. 36 ): phallobase moderately long, PhL/PrL: 1.67 (n=1); parameres weakly curved ventrad, with rounded apices (lateral aspect); penis narrowly rounded at apex; sclerotised fibula present. Ovipositor with right coxite ca 1.3× as long as left coxite; bursa copulatrix with numerous scattered microsclerites ( Fig. 37B ). Secondary sexual dimorphism Male abdominal ventrite 5 narrowly truncate at apex, female ventrite 5 arcuate at apex with short apical keel. Distribution Thailand ( Fig. 110C ).