Gonaphodioides zempoaltepetlensis (Coleoptera: Scarabaeidae: Aphodiinae), a new species from Oaxaca, Mexico Author Dellacasa, Marco Author Dellacasa, Giovanni Author Arriaga-Jiménez, Alfonsina text Zootaxa 2018 2018-11-02 4508 4 587 593 journal article 28028 10.11646/zootaxa.4508.4.9 6b6a2b6e-af31-483d-a17b-8521c988e4ac 1175-5326 2607558 B5173306-FEA0-4FF4-A360-620C4F057EE3 Key to species of Gonaphodioides modified from Dellacasa et al. (2012) 1. Elytra denticulate at humeral angle ................................................................................................................................................... 2 - Elytra not denticulate at humeral angle but therein with distinctly raised epipleural carina ............................................................ 8 2. Elytra subparallel sided, elongate (see Dellacasa et al . 2012 ). Length 5.5–6.0 mm. Bolivia , Colombia ........................................... ....................................................................................................................................................... G. acutecernans (Balthasar, 1960) - Elytra oval, less elongate (see Dellacasa et al . 2012 ). Length 3.5–6.0 mm...................................................................................... 3 3. Elytral interstriae distinctly convex. Length 4.0–5.0 mm. Colombia , Venezuela ................................ G. columbicus (Harold, 1880) - Elytral interstriae flat or almost flat. Length 3.5–6.0 mm................................................................................................................. 4 4. Pronotum dually punctate; few large punctures irregularly scattered on sides; small punctures evenly, sparsely distributed throughout.......................................................................................................................................................................................... 5 - Pronotum simply, evenly, or almost evenly punctate ........................................................................................................................ 6 5. Elytral striae fine, sparsely punctate, faintly crenulate; interstriae flat with extremely fine, sparse punctures. Black. Length 5.0– 6.0 mm. Mexico ( Chiapas ) ................................................................................... G. skelleyi Dellacasa, Dellacasa, & Gordon, 2012 - Elytral striae deep, coarsely, densely punctate, crenulate; interstriae feebly convex, sparsely but distinctly punctate. Piceous black; elytra with yellow spots variously arranged. Length 3.5–4.0 mm. El Salvador ....................................................................... ........................................................................................................................ G. cartwrighti Dellacasa, Dellacasa, & Gordon, 2013 6. Pronotum lateral margins strongly inwardly sinuate before hind angles. Dorsal surface yellowish brown, head anteriorly and sides of pronotum lighter; elytra with juxtasutural interstria, base of interstria 5 and preapical declivity brown, apical declivity with an oblique oval yellow spot extended from interstria 2–7 and with margins dark brown ( Fig. 5 ); sometimes the margins widely faded ( Fig. 6–7 ). Length 4.5–5.0 mm. Mexico ( Oaxaca ) ...................................................................... G. zempoaltepetlensis , new species - Pronotum lateral margins feebly sinuate before hind angles or subparallel. Color uniformly darker brown, occasionally with a vague mark on elytral declivity. Length 3.5–4.5 mm ........................................................................................................................ 7 7. Elytral interstriae coarsely punctate. Dorsal color dark chestnut brown. Length 3.5–4.0 mm Mexico ( Oaxaca ) .............................. ............................................................................................................................ G. ratcliffei Dellacasa, Dellacasa, & Gordon, 2012 - Elytral interstriae finely punctate. Dorsal color dark brown. Length 4.0– 4.5 mm . Mexico ( Hidalgo , Puebla , San Luis Potosí )....... .............................................................................................................................. G. newtoni Dellacasa, Dellacasa, & Gordon, 2012 8. Elytral interstriae flat, superficially microreticulate thus almost dull; sides of pronotum relatively more coarsely and more irregularly punctate. Dorsal color reddish testaceous. Length 3.5–4.0 mm. Bolivia , Brazil , Paraguay ....................................................... ............................................................................................................................................................... G. sincerus (Petrovitz, 1973) - Elytral interstriae convex, smooth thus shiny; sides of pronotum relatively less coarsely and more regularly punctate. Dorsal color chestnut brown. Length 4.0– 4.5 mm . Brazil , Costa Rica , Ecuador , Mexico ( Oaxaca ), Panama , Peru , Venezuela ............................ .................................................................................................................................................................... G. chapini (Hinton, 1934)