Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae) Author De Mattia, Willy https://orcid.org/0000-0002-0056-467X Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria willy.demattia@icgeb.org Author Reier, Susanne Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria Author Haring, Elisabeth Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria text ZooKeys 2021 2021-12-14 1077 1 175 http://dx.doi.org/10.3897/zookeys.1077.67081 journal article http://dx.doi.org/10.3897/zookeys.1077.67081 1313-2970-1077-1 C28AD65A76F242CFBED7DFB3702CABCE 734088641608531C8E2CC69397B000ED Stigmatica paestana intustructa (Westerlund, 1883) Figs 62.4, 62.5, 64.2 Distribution. This taxon is present in Basilicata, known from the area surrounding Balvano (Italy) ( Nordsieck 2013b : 7). Figure 62. Stigmatica paestana paestana (Philippi, 1836), Monte Argentario, Grosseto 62.1 whole distal genital organs 62.2 internal distal part of genital organs 62.3 cross section of the epiphallus. Stigmatica paestana intustructa (Westerlund, 1883), Balvano, Potenza 62.4 whole distal genital organs 62.5 internal distal part of genital organs. Specimens examined. Italy , Basilicata , Potenza , Balvano , near the castle. 450 m asl , 40°39'01.23"N , 15°30'38.91"E , W. De Mattia and J. Macor leg. and det., 3 dissected spm ; Italy , Basilicata , Potenza , Balvano. 450 m asl , 40°38'40.44"N , 15°30'39.09"E , I. Niero leg. and det., 2 dissected spm . External morphology of the genital organs (Fig. 62 .4). The V is much shorter than the FO. The FDBC is longer than the SDBC+BC. The BC+SDBC is slightly club-like in shape, with a more or less clear distinction between the SDBC and the BC. The D is much longer than the SDBC+BC and much thinner. The V is short but wide in diameter. The PC is ~ 4 x longer than the V. The P is cylindrical and slightly swollen. The transition between P and EP is clearly visible with a narrowing and the presence of the ER. The PR is short and robust. The E is longer than the P. Internal morphology of the genital organs (Fig. 62 .5). The V, the A and the P lack any macro-sculpturing. The internal surface shows a coarse fine granulation. The pseudopapilla is elongated and with a smooth surface. It originates directly from the wall of the E. Its basis is connected with one ELP. The E shows two moderately fringed ELP. They proximally fade before the VD. The epiphallar formula is PP(ELP). Remarks. Stigmatica paestana intustructa shows an anatomy of the genital organs very similar to the nominate subspecies.