<document ID-DOI="http://doi.org/10.5281/zenodo.3728763" ID-GBIF-Dataset="5b53b2fd-98cb-4450-96cc-b8e26c1e093f" ID-GBIF-Taxon="162716779" ID-Zenodo-Dep="3728763" approvalRequired="24" approvalRequired_for_document="1" approvalRequired_for_matCits="2" approvalRequired_for_textStreams="21" checkinTime="1584986693040" checkinUser="valdenar" docAuthor="Solis, M. Alma, Rodríguez, Eugenie Phillips-, Hallwachs, Winnie, Dapkey, Tanya &amp; Janzen, Daniel H." docDate="2020" docId="03E087B3DF368A2EFF13A283FEF4FA6C" docLanguage="en" docName="PROC.ENTOMOL.SOC.WASH.122.1.147.pdf.imf" docOrigin="Proceedings of the Entomological Society of Washington 122 (1)" docStyle="DocumentStyle{}" docStyleName="procentomolsocwash.2010.journal_article" docTitle="Asturodes encisoensis Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey 2020" docType="treatment" docVersion="6" lastPageId="21" lastPageNumber="168" masterDocId="FFD9FFCBDF228A3BFFF6A034FFBDFFE5" masterDocTitle="Asturodes Amsel (Lepidoptera: Crambidae: Spilomelinae): Three New Species From The Western Hemisphere And Food Plant Records From Area De Conservación Guanacaste, Costa Rica" masterLastPageNumber="171" masterPageNumber="147" pageId="20" pageNumber="167" updateTime="1643439281101" updateUser="ExternalLinkService">
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<mods:title>Asturodes Amsel (Lepidoptera: Crambidae: Spilomelinae): Three New Species From The Western Hemisphere And Food Plant Records From Area De Conservación Guanacaste, Costa Rica</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solis, M. Alma</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rodríguez, Eugenie Phillips-</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hallwachs, Winnie</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dapkey, Tanya</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janzen, Daniel H.</mods:namePart>
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<mods:title>Proceedings of the Entomological Society of Washington</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>2020-02-29</mods:number>
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<mods:number>122</mods:number>
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<mods:number>1</mods:number>
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<mods:identifier type="DOI">10.4289/0013-8797.122.1.147</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3728763" ID-GBIF-Taxon="162716779" ID-Zenodo-Dep="3728763" LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C" httpUri="http://treatment.plazi.org/id/03E087B3DF368A2EFF13A283FEF4FA6C" lastPageId="21" lastPageNumber="168" pageId="20" pageNumber="167">
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<paragraph blockId="20.[229,599,695,753]" box="[229,599,695,720]" pageId="20" pageNumber="167">
<heading box="[229,599,695,720]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="167" reason="3">
<emphasis bold="true" box="[229,599,695,720]" pageId="20" pageNumber="167">
<taxonomicName authority="Solis" authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[229,594,695,720]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="167" phylum="Arthropoda" rank="species" species="encisoensis" status="new species">
<emphasis bold="true" box="[229,519,695,720]" italics="true" pageId="20" pageNumber="167">Asturodes encisoensis</emphasis>
Solis
</taxonomicName>
,
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="167" type="description">
<paragraph blockId="20.[229,599,695,753]" box="[335,494,728,753]" pageId="20" pageNumber="167">
<heading box="[335,494,728,753]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="167" reason="3">
<emphasis bold="true" box="[335,494,728,753]" pageId="20" pageNumber="167">new species</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[177,651,778,869]" pageId="20" pageNumber="167">
http://zoobank.org/ 
<uuid pageId="20" pageNumber="167">ACB16A3F-6593- 4AED-9150-5526F0AE7136</uuid>
(
<figureCitation box="[257,380,844,869]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 1‒7. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="20" pageNumber="167">Figs. 4, 7</figureCitation>
, 
<figureCitation box="[395,428,844,869]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 8–11. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="20" pageNumber="167">11</figureCitation>
, 
<figureCitation box="[444,524,844,869]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 12–19. Asturodes male genitalia and phallus. 12–13, A. junkoshimurae, USNM slide 115489. 14–15, A. fimbriauralis, USNM slide 115491. 16–17, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="20" pageNumber="167">18–19</figureCitation>
, 
<figureCitation box="[539,572,844,869]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 20–23. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="20" pageNumber="167">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="167" type="diagnosis">
<paragraph blockId="20.[159,670,893,1749]" pageId="20" pageNumber="167">
Diagnosis.—Forewing with a dark brown line on M2 near margin, only a spot on M1 with no dark brown bar to reniform spot, reniform spot a dark brown “upside down U” over a “right-side up U” (
<figureCitation box="[580,664,1026,1051]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 1‒7. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="20" pageNumber="167">Figs. 4</figureCitation>
, 
<figureCitation box="[159,191,1059,1084]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 8–11. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="20" pageNumber="167">11</figureCitation>
, see red circle); male genitalia with uncus tip pointed (
<figureCitation box="[394,487,1093,1118]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 12–19. Asturodes male genitalia and phallus. 12–13, A. junkoshimurae, USNM slide 115489. 14–15, A. fimbriauralis, USNM slide 115491. 16–17, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="20" pageNumber="167">Fig. 18</figureCitation>
); female genitalia with signa on corpus bursae in two dorsal longitudinal lines (
<figureCitation box="[473,565,1159,1184]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 20–23. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="20" pageNumber="167">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="167" type="description">
<paragraph blockId="20.[159,670,893,1749]" lastBlockId="20.[701,1212,196,1749]" pageId="20" pageNumber="167">
Description.—Adult. 
<emphasis box="[485,551,1193,1217]" italics="true" pageId="20" pageNumber="167">Head</emphasis>
: Scape posteriorly with a few dark brown scales, vertex only with yellow scales. Labial palpus (
<figureCitation box="[258,332,1292,1317]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 1‒7. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="20" pageNumber="167">Fig. 7</figureCitation>
) with first and second segments yellow and with a dark brown band anteriorly, only dorsally, third segment minute, dark brown on male, yellow on female. Proboscis with yellow scales throughout, without dark brown scales as proboscis curves under. 
<emphasis box="[460,559,1491,1515]" italics="true" pageId="20" pageNumber="167">Thorax:</emphasis>
Average forewing length 
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(n = 8) in males, 
<quantity box="[159,276,1558,1583]" metricMagnitude="-3" metricUnit="m" metricValue="8.42" pageId="20" pageNumber="167" unit="mm" value="8.42">8.42 mm</quantity>
(n = 7) in females. Forewing pattern as in 
<figureCitation box="[338,460,1591,1616]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 1‒7. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="20" pageNumber="167">Figures 4</figureCitation>
and 
<figureCitation box="[533,566,1591,1616]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 8–11. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="20" pageNumber="167">11</figureCitation>
, a dark brown line posterior to M2 near margin, only a spot on 
<botanicCollectionCode box="[335,362,1658,1681]" pageId="20" pageNumber="167">M</botanicCollectionCode>
1 with no dark brown line to reniform spot, reniform spot a dark brown “upside down 
<botanicCollectionCode box="[417,441,1724,1749]" pageId="20" pageNumber="167">U</botanicCollectionCode>
” over a “right-side up 
<botanicCollectionCode box="[744,768,196,221]" pageId="20" pageNumber="167">U</botanicCollectionCode>
” (
<figureCitation box="[801,896,196,221]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 8–11. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="20" pageNumber="167">Fig. 11</figureCitation>
, see red circle); no dark brown markings in radial area between reniform spot and postmedial line; first lengthwise line of zigzag between Rs3 and Rs4 shortened, almost a spot; second lengthwise line of zigzag between Rs4 and 
<botanicCollectionCode box="[753,781,396,419]" pageId="20" pageNumber="167">M</botanicCollectionCode>
1 very short, a spot; antemedial line incomplete. 
<emphasis box="[860,976,429,453]" italics="true" pageId="20" pageNumber="167">Abdomen</emphasis>
: Third abdominal tergite with anteromedial dark spot and two lateral triangles, fourth tergite with anteromedial dark spot, fifth tergite with silver transverse band posteriorly (
<figureCitation box="[1122,1196,561,586]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 1‒7. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="20" pageNumber="167">Fig. 4</figureCitation>
). Male genitalia (
<figureCitation box="[902,1053,595,620]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 12–19. Asturodes male genitalia and phallus. 12–13, A. junkoshimurae, USNM slide 115489. 14–15, A. fimbriauralis, USNM slide 115491. 16–17, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="20" pageNumber="167">Figs. 18–19</figureCitation>
) uncus with very short, lateral setae; uncus &quot;tip&quot; pointed and prominent. Parateguminal androconia oblong, broader anteriorly. Valva with fibula, or sclerotized, fingerlike process extending only slightly beyond saccular margin. Juxta medially with a lightly sclerotized, single line. Phallus vesica with large spinules. Female genitalia (
<figureCitation box="[822,910,893,918]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 20–23. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="20" pageNumber="167">Fig. 23</figureCitation>
) with ductus bursae with a square-like antrum about 
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in width, followed by two elongate, sclerotized structures within that are as wide as the band, strongly constricted and then ductus bursae three times as wide as antrum forming a lightly sclerotized pouchlike structure, then constricted at entrance to corpus bursae. Corpus bursae round, signum two longitudinal lines, half the length of corpus bursae. Ductus seminalis originating shortly after antrum.
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="168" pageId="20" pageNumber="167" type="etymology">
<paragraph blockId="20.[701,1212,196,1749]" pageId="20" pageNumber="167">
Etymology.—The species epithet “ 
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="167" phylum="Arthropoda" rank="species" species="encisoensis">
<emphasis italics="true" pageId="20" pageNumber="167">encisoensis</emphasis>
</taxonomicName>
” is a noun derived from the name of the 
<typeStatus box="[869,923,1359,1383]" pageId="20" pageNumber="167">type</typeStatus>
locality, Teniente Agripino Enciso National Park in 
<collectingCountry box="[1091,1207,1393,1417]" name="Paraguay" pageId="20" pageNumber="167">Paraguay</collectingCountry>
, and “ 
<emphasis box="[767,828,1426,1449]" italics="true" pageId="20" pageNumber="167">ensis</emphasis>
,” a Latin suffix meaning place. Distribution.—This species is found in one of the protected natural areas of the Paraguayan Chaco in the Teniente (Lieutenant) Agripino Enciso National Park in northwestern 
<collectingCountry box="[994,1112,1592,1616]" name="Paraguay" pageId="20" pageNumber="167">Paraguay</collectingCountry>
, in the Argentinian 
<collectingRegion box="[871,951,1624,1649]" country="Argentina" name="Chaco" pageId="20" pageNumber="167">Chaco</collectingRegion>
in Las Brenas in northern 
<collectingCountry box="[831,960,1657,1682]" name="Argentina" pageId="20" pageNumber="167">Argentina</collectingCountry>
, and in northern 
<collectingCountry box="[701,831,1690,1715]" name="Venezuela" pageId="20" pageNumber="167">Venezuela</collectingCountry>
just to the west and outside of Parque Nacional Aguaro-Guariquito.
</paragraph>
<paragraph blockId="21.[109,619,196,1417]" pageId="21" pageNumber="168">Biological remarks.—Adults are attracted to lights. Nothing is known about the larval biology of this species.</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="168" type="materials_examined">
<paragraph blockId="21.[109,619,196,1417]" pageId="21" pageNumber="168">
<typeStatus box="[140,202,297,321]" pageId="21" pageNumber="168">Type</typeStatus>
Material.— 
<materialsCitation ID-GBIF-Occurrence="2579594601" collectingDate="1926-03-28" collectionCode="USN-" collectorName="M. Pogue &amp; M. A. Solis" location="Nueva" municipality="Dep" pageId="21" pageNumber="168" specimenCode="MENT01116302" specimenCount="1" specimenCount-male="1" stateProvince="PARAGUAY" typeStatus="holotype">
<typeStatus box="[382,550,296,321]" pageId="21" pageNumber="168">HOLOTYPE</typeStatus>
<specimenCount box="[587,610,295,320]" pageId="21" pageNumber="168" type="male">♂</specimenCount>
: 
<collectingRegion box="[109,272,329,354]" country="Paraguay" name="Asuncion" pageId="21" pageNumber="168">PARAGUAY</collectingRegion>
: 
<collectingMunicipality box="[312,364,330,353]" pageId="21" pageNumber="168">Dep</collectingMunicipality>
[ar]tamento 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFDEBA17EFDD7FE84" box="[541,618,330,353]" municipality="Dep" name="Nueva" pageId="21" pageNumber="168" stateProvince="PARAGUAY">Nueva</location>
<collectingRegion box="[109,227,362,387]" country="Paraguay" name="Asuncion" pageId="21" pageNumber="168">Asuncion</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFF05A15FFF1DFE41" municipality="Dep" name="Parque Nacional Teniente Enciso" pageId="21" pageNumber="168" stateProvince="PARAGUAY">Parque Nacional Teniente Enciso</location>
, 
<geoCoordinate box="[179,292,395,420]" pageId="21" pageNumber="168">
61 
<emphasis box="[209,221,395,420]" italics="true" pageId="21" pageNumber="168">°</emphasis>
39 
<emphasis bold="true" box="[252,262,395,420]" italics="true" pageId="21" pageNumber="168">̍</emphasis>
W
</geoCoordinate>
, 
<geoCoordinate box="[312,413,395,420]" pageId="21" pageNumber="168">
21 
<emphasis box="[342,354,395,420]" italics="true" pageId="21" pageNumber="168">°</emphasis>
13 
<emphasis bold="true" box="[384,394,395,420]" italics="true" pageId="21" pageNumber="168">̍</emphasis>
S
</geoCoordinate>
, 
<collectorName box="[434,560,396,420]" pageId="21" pageNumber="168">M. Pogue</collectorName>
and 
<collectorName box="[109,246,428,453]" pageId="21" pageNumber="168">M.A. Solis</collectorName>
, 
<date box="[272,440,428,453]" pageId="21" pageNumber="168" value="1926-03-28">
<collectingDate box="[272,440,428,453]" pageId="21" pageNumber="168" value="1926-03-28">26‒28 March</collectingDate>
</date>
, 1986 [
<collectionCode box="[550,616,428,453]" pageId="21" pageNumber="168">USN-</collectionCode>
<specimenCode box="[109,303,462,487]" pageId="21" pageNumber="168">MENT01116302</specimenCode>
].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594624" pageId="21" pageNumber="168" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus box="[327,486,462,487]" pageId="21" pageNumber="168">PARATYPES</typeStatus>
: 
<specimenCount box="[504,538,461,487]" pageId="21" pageNumber="168" type="male">1♂</specimenCount>
, same data as 
<typeStatus box="[229,324,495,520]" pageId="21" pageNumber="168">holotype</typeStatus>
[USNMENT01116303].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594595" collectingDate="1940-12-18" collectorName="H. L. Parker" country="Argentina" location="Las Brenas" municipality="Las Brenas" pageId="21" pageNumber="168" specimenCount="1" specimenCount-male="1" stateProvince="Chaco">
<collectingCountry box="[109,296,528,553]" name="Argentina" pageId="21" pageNumber="168">ARGENTINA</collectingCountry>
: 
<collectingRegion box="[310,393,528,553]" country="Argentina" name="Chaco" pageId="21" pageNumber="168">Chaco</collectingRegion>
, 
<specimenCount box="[409,448,527,553]" pageId="21" pageNumber="168" type="male">1♂</specimenCount>
, 
<collectingMunicipality box="[467,614,529,552]" pageId="21" pageNumber="168">Las Brenas</collectingMunicipality>
, Lot No. 
<geoCoordinate box="[235,359,561,586]" pageId="21" pageNumber="168">40-25771</geoCoordinate>
, 
<date box="[382,537,561,586]" pageId="21" pageNumber="168" value="1940-12-18">
<collectingDate box="[382,537,561,586]" pageId="21" pageNumber="168" value="1940-12-18">18.XII.1940</collectingDate>
</date>
, 
<collectorName pageId="21" pageNumber="168">H.L. Parker</collectorName>
, [USNMENT01116300].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594622" collectingDate="1988-05-25" collectionCode="USNM" collectorName="M. Epstein &amp; C. Canaday" latitude="8.57" location="Hato Masaguarat" longLatPrecision="781" longitude="-67.58" municipality="Guarico" pageId="21" pageNumber="168" specimenCount="2" specimenCount-female="2">
VEN- EZUELA: 
<collectingMunicipality box="[257,363,628,653]" pageId="21" pageNumber="168">Guarico</collectingMunicipality>
: 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFE76A241FDDAFD68" box="[384,615,629,653]" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Guarico" name="Hato Masaguarat" pageId="21" pageNumber="168">Hato Masaguarat</location>
: 
<specimenCount box="[109,170,660,686]" pageId="21" pageNumber="168" type="female">2♀♀</specimenCount>
, 
<locationDeviation bearing="180.0" box="[185,286,661,686]" distance="45000.0" pageId="21" pageNumber="168">
<quantity box="[185,263,661,686]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="168" unit="km" value="45.0">45 km</quantity>
S
</locationDeviation>
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFED3A2A1FE22FD4B" box="[293,415,661,686]" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Guarico" name="Calabozo" pageId="21" pageNumber="168">Calabozo</location>
, 
<geoCoordinate box="[426,527,661,686]" degrees="67.58" direction="west" orientation="longitude" pageId="21" pageNumber="168" precision="555" value="-67.58">67.58W</geoCoordinate>
<geoCoordinate box="[533,614,661,686]" degrees="8.57" direction="north" orientation="latitude" pageId="21" pageNumber="168" precision="555" value="8.57">8.57N</geoCoordinate>
, 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFF9BA282FF20FD2A" box="[109,157,694,719]" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Guarico" name="Gal" pageId="21" pageNumber="168">Gal</location>
[le]ry 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFF05A283FEF9FD2B" box="[243,324,695,718]" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Guarico" name="Forest" pageId="21" pageNumber="168">Forest</location>
, uv light, 
<date box="[482,613,694,719]" pageId="21" pageNumber="168" value="1988-05-25">
<collectingDate box="[482,613,694,719]" pageId="21" pageNumber="168" value="1988-05-25">25.V.1988</collectingDate>
</date>
, 
<collectorName box="[109,258,727,752]" pageId="21" pageNumber="168">M. Epstein</collectorName>
and 
<collectorName box="[344,504,727,752]" pageId="21" pageNumber="168">C. Canaday</collectorName>
, 
<collectionCode box="[528,618,727,752]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="168" type="Museum">USNM</collectionCode>
slide #115503 [USNMENT01116288, USNMENT01116293];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594618" collectingDate="1988-05-13" collectingDateMax="1988-05-16" collectingDateMin="1988-05-13" collectionCode="USNM, USN-" collectorName="M. Epstein &amp; R." location="Blahnik" pageId="21" pageNumber="168" specimenCode="MENT01116290" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount box="[429,467,793,819]" pageId="21" pageNumber="168" type="male">1♂</specimenCount>
, 
<specimenCount box="[499,538,793,819]" pageId="21" pageNumber="168" type="female">1♀</specimenCount>
, 
<date pageId="21" pageNumber="168" value="1988-05-13" valueMax="1988-05-16" valueMin="1988-05-13">
<collectingDate pageId="21" pageNumber="168" value="1988-05-13" valueMax="1988-05-16" valueMin="1988-05-13">13‒ 16.V.1988</collectingDate>
</date>
, 
<collectorName box="[259,402,827,852]" pageId="21" pageNumber="168">M. Epstein</collectorName>
and 
<collectorName box="[476,505,828,851]" pageId="21" pageNumber="168">R.</collectorName>
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFDF1A30FFDD6FCB1" box="[519,619,827,852]" name="Blahnik" pageId="21" pageNumber="168">Blahnik</location>
<collectionCode box="[109,199,860,885]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="168" type="Museum">USNM</collectionCode>
slide #115504, #115501 [
<collectionCode box="[542,618,860,885]" pageId="21" pageNumber="168">USN-</collectionCode>
<specimenCode box="[109,327,893,918]" pageId="21" pageNumber="168">MENT01116290</specimenCode>
, USNMENT0111 6294];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594611" collectingDate="1988-05-20" collectionCode="USNM" collectorName="M. Epstein &amp; R." location="Blahnik" pageId="21" pageNumber="168" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount box="[204,243,926,952]" pageId="21" pageNumber="168" type="male">1♂</specimenCount>
, 
<specimenCount box="[265,303,926,952]" pageId="21" pageNumber="168" type="female">1♀</specimenCount>
, 
<date box="[325,456,927,952]" pageId="21" pageNumber="168" value="1988-05-20">
<collectingDate box="[325,456,927,952]" pageId="21" pageNumber="168" value="1988-05-20">20.V.1988</collectingDate>
</date>
, 
<collectorName box="[475,618,927,952]" pageId="21" pageNumber="168">M. Epstein</collectorName>
and 
<collectorName box="[168,196,961,984]" pageId="21" pageNumber="168">R.</collectorName>
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFF24A3F4FE84FC3C" box="[210,313,960,985]" name="Blahnik" pageId="21" pageNumber="168">Blahnik</location>
, 
<collectionCode box="[332,422,960,985]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="168" type="Museum">USNM</collectionCode>
slide #115639 [USNMENT01116296;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594623" pageId="21" pageNumber="168" specimenCount="1">USNMENT0- 1116295];</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594625" collectingDate="1988-06-03" collectingDateMax="1988-06-05" collectingDateMin="1988-06-03" collectionCode="USN-" collectorName="M. Epstein" pageId="21" pageNumber="168" specimenCode="MENT01116292" specimenCount="2" specimenCount-male="2">
<specimenCount box="[253,315,1026,1051]" pageId="21" pageNumber="168" type="male">2♂♂</specimenCount>
, 
<date box="[338,502,1026,1051]" pageId="21" pageNumber="168" value="1988-06-03" valueMax="1988-06-05" valueMin="1988-06-03">
<collectingDate box="[338,502,1026,1051]" pageId="21" pageNumber="168" value="1988-06-03" valueMax="1988-06-05" valueMin="1988-06-03">3‒5.VI.1988</collectingDate>
</date>
, 
<collectorName pageId="21" pageNumber="168">M. Epstein</collectorName>
[USNMENT01116291, 
<collectionCode box="[548,618,1059,1084]" pageId="21" pageNumber="168">USN-</collectionCode>
<specimenCode box="[109,320,1093,1118]" pageId="21" pageNumber="168">MENT01116292</specimenCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594616" collectingDate="1989-07-05" collectingDateMax="1989-07-07" collectingDateMin="1989-07-05" collectorName="M. Epstein" pageId="21" pageNumber="168" specimenCount="1" specimenCount-female="1">
<specimenCount box="[344,381,1092,1118]" pageId="21" pageNumber="168" type="female">1♀</specimenCount>
, 
<date box="[402,566,1093,1118]" pageId="21" pageNumber="168" value="1989-07-05" valueMax="1989-07-07" valueMin="1989-07-05">
<collectingDate box="[402,566,1093,1118]" pageId="21" pageNumber="168" value="1989-07-05" valueMax="1989-07-07" valueMin="1989-07-05">5‒7.VII.1989</collectingDate>
</date>
, 
<collectorName pageId="21" pageNumber="168">M. Epstein</collectorName>
[USNMENT01116297];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594598" collectingDate="1989-07-13" collectingDateMax="1989-07-14" collectingDateMin="1989-07-13" collectionCode="USNM" collectorName="M. Epstein" pageId="21" pageNumber="168" specimenCount="1" specimenCount-female="1">
<specimenCount box="[572,610,1125,1151]" pageId="21" pageNumber="168" type="female">1♀</specimenCount>
, 
<date box="[109,314,1159,1184]" pageId="21" pageNumber="168" value="1989-07-13" valueMax="1989-07-14" valueMin="1989-07-13">
<collectingDate box="[109,314,1159,1184]" pageId="21" pageNumber="168" value="1989-07-13" valueMax="1989-07-14" valueMin="1989-07-13">13‒14.VII.1989</collectingDate>
</date>
, 
<collectorName box="[342,500,1159,1184]" pageId="21" pageNumber="168">M. Epstein</collectorName>
, 
<collectionCode box="[528,618,1159,1184]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="168" type="Museum">USNM</collectionCode>
slide #115638 [USNMENT01116301];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594649" collectingDate="1990-09-20" collectingDateMax="1990-09-21" collectingDateMin="1990-09-20" collectionCode="J" collectorName="M. Epstein" location="Wilterding" pageId="21" pageNumber="168" specimenCount="1" specimenCount-female="1">
<specimenCount box="[109,147,1225,1250]" pageId="21" pageNumber="168" type="female">1♀</specimenCount>
, 
<date box="[169,363,1225,1250]" pageId="21" pageNumber="168" value="1990-09-20" valueMax="1990-09-21" valueMin="1990-09-20">
<collectingDate box="[169,363,1225,1250]" pageId="21" pageNumber="168" value="1990-09-20" valueMax="1990-09-21" valueMin="1990-09-20">20‒21.IX.1990</collectingDate>
</date>
, 
<collectorName box="[382,526,1225,1250]" pageId="21" pageNumber="168">M. Epstein</collectorName>
and 
<collectionCode box="[599,614,1226,1249]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14647" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="21" pageNumber="168" type="Herbarium">J</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFF9BA4DFFF48FAE1" box="[109,245,1259,1284]" name="Wilterding" pageId="21" pageNumber="168">Wilterding</location>
III [USNMENT01116298]; [
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2579594615" collectingDate="1989-07-05" collectingDateMax="1989-07-07" collectingDateMin="1989-07-05" collectionCode="M, USNM, USN-" collectorName="M. Epstein" country="Indonesia" location="Deza" municipality="Morades" pageId="21" pageNumber="168" specimenCode="MENT01116299" specimenCount="1" specimenCount-male="1">
Hato 
<collectingCountry box="[190,264,1292,1317]" name="Indonesia" pageId="21" pageNumber="168">Flores</collectingCountry>
<collectingMunicipality box="[277,384,1292,1317]" pageId="21" pageNumber="168">Morades</collectingMunicipality>
, but same data as above], 
<specimenCount box="[204,241,1324,1350]" pageId="21" pageNumber="168" type="male">1♂</specimenCount>
, 
<date box="[258,420,1325,1350]" pageId="21" pageNumber="168" value="1989-07-05" valueMax="1989-07-07" valueMin="1989-07-05">
<collectingDate box="[258,420,1325,1350]" pageId="21" pageNumber="168" value="1989-07-05" valueMax="1989-07-07" valueMin="1989-07-05">5‒7.VII.1989</collectingDate>
</date>
, 
<collectorName box="[434,566,1325,1350]" pageId="21" pageNumber="168">M. Epstein</collectorName>
and 
<collectionCode box="[109,140,1359,1382]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M�nchen" pageId="21" pageNumber="168" type="Herbarium">M</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:03E087B3DF368A2EFF13A283FEF4FA6C:8E96607EDF378A2EFF69A57BFF62FA83" box="[159,223,1359,1382]" country="Indonesia" municipality="Morades" name="Deza" pageId="21" pageNumber="168">Deza</location>
, 
<collectionCode box="[244,331,1358,1383]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="168" type="Museum">USNM</collectionCode>
slide #115502 [
<collectionCode box="[546,618,1358,1383]" pageId="21" pageNumber="168">USN-</collectionCode>
<specimenCode box="[109,319,1392,1417]" pageId="21" pageNumber="168">MENT01116299</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>