New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae) Author Brown, John W. text Zootaxa 2019 2019-09-17 4671 2 195 222 journal article 25519 10.11646/zootaxa.4671.2.2 978ec4b3-8d0e-4f47-8084-610b34b96759 1175-5326 3442344 8DA2FA3F-3629-4D10-92B0-671637D91DD4 Aethes triassumenta Pogue , new species Figs. 15, 16 , 31 , 45 Diagnosis . Adults of A. triassumenta ( Figs. 15, 16 ) are superficially extremely similar to those of A. bomonana ( Kearfott, 1907 ) and A. obliquana ( Kearfott, 1907 ) , with a slightly shiny, pale cream forewing ground color and illdefined, contrasting brown to red-brown markings, often with numerous tiny brown specks in the apical and subapical regions. The male genitalia of A . triassumenta ( Fig. 31 ) are distinguished from other congeners by the absence of the sickle-shaped processes of the socii and the attachment of the juxta to the valva in the male genitalia; and the extremely elongate, frail, membranous corpus bursae in the female genitalia ( Fig. 45 ). The shape of the valva is not particular divergent from that of other congeners, with a weakly concave outer margin and small lobe or spine near the apex of the costa. The suberect, elongate-triangular socii of A. triassumenta are nearly identical to those of A. tuxtlana , and likely are homologous with the sickle-shaped socii of other Aethes . The species is assigned to Aethes on the basis of the facies and DNA barcode data ( Brown et al . 2019 ). Description . Head: Frons and vertex white to cream; labial palpus reddish brown on outer surface, white to cream on dorsal and inner surface. Thorax: Mesonotum and tegula white with variable cream scaling; posterior crest white; forewing length 8.9–9.1 mm (mean = 9.0 mm, n = 10); forewing ground color white; base of costa and small irregular patch above R vein reddish brown; an irregular horizontal patch extending from wing base to middle of discal cell, consisting of a posterior chestnut patch proximally contiguous with a larger anterior chestnut patch, with interior scaling shading to light cinnamon; two median patches at end of discal cell: anterior one chestnut, usually larger than posterior patch, posterior patch oblong with anterior 0.5 chestnut and posterior 0.5 buff yellow with a few tawny scales proximally; two buff-yellow patches: one from base of R vein to just before horizontal bar below Sc, second one less distinct, from base of R 1 to end of discal cell; both patches below costa; costa and hind margin with short brown striations, variable in number and intensity; subapical band from costa to hind margin white. Fringe white. Hindwing pale gray. Fringe white with basal band of ground color. Abdomen: Unmodified. White to cream with scattered pale-gray scales; genital tuft white. Male genitalia ( Fig. 31 ) with uncus absent; socius with basal 0.25 fused to tegumen, distal part free, narrow-triangular, with pointed apex, setose; transtilla broad at base, with pointed spines laterally, median projection elongate, attenuate distally, rounded apically; valva broad, length about 1.4 times maximum width, costa weakly concave mesially, apically produced into thumblike process with variably developed thorn, sacculus undifferentiated; vinculum arms free distally; juxta shieldlike with prominent lateral arms confluent with upper edge of sacculus. Phallus large, 1.2 times length of valva, ventral projection elongate, curved, pointed; vesica with a single large cornutus. Female genitalia ( Fig. 45 ) with papillae anales slender, lightly sclerotized; apophyses anteriores about 1.2 times length of apophyses posteriores; sterigma mostly membranous, lightly sclerotized around ositum bursae; lamella postvaginalis comprised of two semicircular plates joined mesially; colliculum sclerotized with a ventrally produced digitate process opposite of origin of accessory bursa, and second digitate process near anterior end of colliculum immediately anterad of cup-shaped concavity; ductus bursae elongate, membranous anterad of colliculum, gradually broadened, lacking distinct junction with corpus bursae; corpus bursae extremely elongate, slender, membranous, without spiculae or signum. Types . Holotype ³, USA , Florida , Highlands Co. , Archbold Biological Station , 25 Dec 1957 , R. W. Pease , Jr. ( USNM ) . Paratypes (47³, 3♀ ). USA : FLORIDA : Same locality as holotype , 23 Dec 1957 , 26 Jan 1958 , J. R. Mal- loch (4³) ( USNM ) . Pinellas Co. : St. Petersburg , 31 Jan 1958 , 11 Feb 1958 , R. W. Pease (2³) ( USNM ) . St Johns Co. : Hastings , [no date], [no collector] (2³) ( USNM ) . Volusia Co. : Cassadaga , 19 Jan 1962 (1³), S. V. Fuller ( USNM ) . Cassadaga , 24 Feb 1960 ( 1♀ ), S. W. Frost ( USNM ) . NORTH CAROLINA : Brunswick Co. : Meyers Clemmins Tract , Green Swamp , 25 Mar 1994 (1³), Sullivan et al . ( USNM ) . Green Swamp , Big Island Annex, 19 Sep 1994 (3³), Sullivan et al . ( USNM ) . MOTSU , inter. firebreaks 32 & 34, 9 May 1994 (1³), 29 Aug 1994 (1³), Sullivan et al . ( USNM ) . Carteret Co. : Willis Road Savannah , 18 Jul 1983 , 24 Aug 1983 , 27 Aug 1983 , 9 Sep 1983 , 20 Sep 1983 , J. B. Sullivan (10³) ( USNM ) . Craven Co. : Rd 122-2, powerline sav., Croatan National Forest , 7 Sep 1999 (1³), J. B. Sullivan ( USNM ) . Croatan Road 147, 2 June 1997 ( 1♀ ), J. B. Sullivan ( USNM ) . New Hanover Co. : MOTSU buffer, Peter’s Point , 27 Sep 1994 (1³), 29 Aug 1994 (1³), Sullivan et al . ( USNM ) . Carteret Co. : Sam Hatcher Road , N34 44.719 , W076 55.659 , Croatan National Forest , 22 Jun 2008 (1³), J. B. Sullivan ( USNM ) . Camp Sam Hatcher Road , 29 Mar 2002 (5³), J. B. Sullivan ( USNM ) . Old landfill off Hobbs Road , 29 Mar 2002 (4³, 1♀ ), J. B. Sullivan ( USNM ) . Willis Savanna , 14 Mar 2002 (1³), J. B. Sullivan ( USNM ) . North of Millis Sa- vanna, Croatan National Forest , 14 Mar 2002 (3³), J. B. Sullivan ( USNM ) . Jct state hiway 101 & 181, 8 Aug 1972 (1³), J. B. Sullivan ( USNM ) . Onslow Co. : Camp Lejune Marine Base , 9 Apr 1999 (1³), J. B. Sullivan ( USNM ) . Pender Co. : Holly shelter gamelands, 12 Apr 1995 , 10 June 1995 , J. B. Sullivan (3³, 1♀ ) ( USNM ) . Distribution and biology . Aethes triassumenta is apparently restricted to the southeastern U.S. , ranging from Florida to North Carolina . Three specimens from Florida were misidentified by Kearfott (1907) and included in his type series of Aethes obliquana . Aethes triassumenta is recorded sporadically throughout the year with no conspicuous peak fight period(s). The early stages are unknown. Remarks . Pogue’s (1986) manuscript name triassumenta was proposed as the type species of his manuscript genus Morta . However, because triassumenta appears to fit rather convincingly in Aethes based on facies and genital morphology, the nomen nudum Morta Pogue and Mickevitch, 1990 is placed under Aethes .